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of the
FLORIDA STATE MUSEUM
Biological Sciences
Volume 23 1978 Number 1
TYPE CATALOGUE OF THE NORTH AMERICAN
CYPRINID FISH GENUS NOTROPIS
CARTER R. GILBERT
UNIVERSITY OF FLORIDA
GAINESVILLE
Numbers of the BULLETIN OF THE FLORIDA STATE MUSEUM, BIOLOGICAL SCIENCES,
are published at irregular intervals. Volumes contain about 300 pages and are not necessarily
completed in any one calendar year.
OLIVER L. AUSTIN, Jr., Editor
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Consultants for this issue:
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ROBERT RUSH MILLER
Communications concerning purchase or exchange of the publications and all manuscripts
should be addressed to: Managing Editor, Bulletin; Florida State Museum; University of Florida:
Gainesville, Florida 32611.
Publication date: June 23, 1978
This public document was promulgated at an annual cost of $4,866.02 or
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terested persons the results of researches in the natural sciences, empha-
sizing the circum-Caribbean region.
Price: $5.00
TYPE CATALOGUE OF THE NORTH AMERICAN CYPRINID
FISH GENUS NOTROPIS
CARTER R. GILBERT'
SYNOPSIS: A type catalogue is presented for the North American cyprinid fish genus Notropis.
This genus, as presently constituted, is the largest of any group of North American fishes, com-
prising 131 valid described species and subspecies (119 and 12, respectively, of which one of the
species is a fossil), and at least a dozen forms that are still undescribed. In addition, seven described
species presently included in the genus Hybopsis may ultimately be transferred to Notropis. A
total of 304 different names are available for the 138 valid described forms.
All nominal forms are listed alphabetically by species or subspecies name. The following
data appear under each separate heading: (1) original genus and species combination, together
with describer and year of description; (2) abbreviated citation of original description; (3) type
locality; (4) catalogue numbers) of holotype, lectotype, neotype, or syntypes, together with
numbers of specimens, range in standard length in mm, and name of collector and date of col-
lection, if known; (5) catalogue numbers) of paratypes, number of specimens in each series, and
an indication of which are paratopotypes; (6) remarks concerning the type (counts and con-
dition), nomenclatural history of the taxon, and definitive references; (7) exact date of original
description if known, or, if not known, the most accurate date determinable; and (8) present
status of the taxon.
In addition to the above list, the following supplementary lists are included: (1) a generic
synonymy; (2) presently recognized species of Notropis, with their included subspecies (if any)
and junior and senior synonyms (if any); (3) species presently referred to the genus Hybopsis,
hut whose affinities are with Notropis and in which genus they may eventually be placed;
(4) nominal species of Notropis that are based on hybrids, one or both parents of which belong
to that genus; (5) nominal species of Notropis that are based on hybrids, both parents of which
belong to other genera; (6) nominal nuda with their identifications; and (7) species that were
originally described in Notropis, but which are now referred to another genus. Also, the genus
is defined and a brief discussion of the nomenclatural history of the group is presented, which
includes a comparison of the numbers of forms described during each decade.
TABLE OF CONTENTS
IN TRODUCTION .. .......... ....... .... ..... ........ ..... 2
DEFINITION OF GENUS Notropis... .. ........ ...................... 11
TAXONOM IC H ISTORY OF G ENUS Notropis .............. ........................................... ............ ...... 12
G EN ERIC SYNO N YM Y .. . ... ... .. ... ...... ................................................... .... 15
STATUS OF G ENUS N otropis .. .... .. .................... .......................................... .......... ........ 16
GENERA THAT MAY ULTIMATELY BE INCLUDED IN Notropis ........... ...................... 17
TAXONOM IC L ISTS.. ... .... ............................. ...... 17
'The author is Associate Curator in Ichthyology at the Florida State Museum and Associate Professor in the Department of
Zixilogy, University of Florida, Gainesville, Florida 32611.
GILBERT, CARTER R. 1978. Type Catalogue of the North American Cyprinid Fish Genus
Notropis. Bull. Florida State Mus., Biol. Sci. 23(1): 1-104.
BULLETIN FLORIDA STATE MUSEUM
VALID FORMS OF Notropis, INCLUDING SYNONYMS ............................. 17
VALID FORMS OF Hybopsis, INCLUDING SYNONYMS ....... .................... ......... .... 21
NOMINAL FORMS OF Notropis BASED ON HYBRIDS OF THE GENUS ....................... ... 21
NOMINAL FORMS OF Notropis BASED ON HYBRIDS OF OTHER GENERA................. .....22
N om ina nuda OF N otropis ............... ...... ...... .......... .................... 22
NOMINAL FORMS OF Notropis Now REFERRED TO OTHER GENERA ............................. 22
ACCOUNTS OF FORMS REFERRED TO NotropiS ........... ...................................... 22
ACCOUNTS OF FORMS REFERRED TO Hybopsis ................. .. .............................. 90
L ITE RA T U R E C IT E D ............ .. ............................................ ........ ..... ... ......... ............. ...... 93
INTRODUCTION
In any systematic study, considerable time and effort is expended in lo-
cating and examining type material and in determining to which species the
many nominal forms refer. This task can be greatly alleviated by previously
published compilations of type material. Such catalogues exist, in published
form, for only five major North American fish collections: the Field Museum
of Natural History (Grey 1947), the Illinois Natural History Survey (Smith
and Bridges 1960), the National Museum of Canada (McAllister 1965), Tulane
University (Suttkus 1970), and the Florida State Museum (Gilbert 1974). In
addition, Henn (1928) and B6hlke (1953) published fish-type catalogues for
the Carnegie Museum and Stanford University, respectively. The Carnegie
Museum's fish collection was sold to the Field Museum of Natural History in
June, 1951 (Loren P. Woods, in litt.), and the Stanford University holdings
were recently incorporated into the collection of the California Academy of
Sciences; Henn's and Bohlke's papers thus become partial catalogues for the
Field Museum and California Academy collections. Of equal value are type
catalogues dealing with specific groups of fishes, which draw together such
information as primary synonymies for the different species, extant type ma-
terial and its disposition, composition of syntypic series, and pertinent infor-
mation on the types themselves. So far, the only compilation of this kind for
any group of North American fishes (freshwater or marine) is that by Collette
and Knapp (1967) for the darters (family Percidae). In no other group of North
American freshwater fishes is the need for such a catalogue more acute than
for the Cyprinidae. Many species in this family, particularly those of the east-
ern genus Notropis, are very similar morphologically, and this, together with
the large number of taxa, has resulted in considerable confusion regarding the
identity of many nominal forms. Several species have been described as new a
number of times. For example, two of the most common and widespread
species, Notropis cornutus (Mitchill 1817) and Notropis lutrensis (Baird and
Girard 1853), have totals of 14 and 25 junior synonyms, respectively, although
at least four of the latter are considered to be valid subspecies (Contreras
1975). Over 625 specific and subspecific names exist for the nearly 300 valid
described forms of North American Cyprinidae (another 20 to 25 forms re-
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
main to be described), of which 295 apply to the 131 valid described species
and subspecies presently included in the genus Notropis. Further complicating
the picture is the fact that many North American cyprinids are known to hy-
bridize, and some of these hybrid combinations have been described as new
species.
The present catalogue of the cyprinid genus Notropis was compiled in
order to answer some of the above needs. Hopefully, this will stimulate the
publication of other such catalogues, so that eventually all genera of North
American Cyprinidae, and perhaps other families as well, will be so treated.
With few exceptions the type of information presented is identical to that
given by Collette and Knapp (1967) for the Percidae. I have attempted to ex-
amine, or at least to verify, the location of extant type material holotypess,
lectotypes, neotypes, syntypes, lectoparatypes, and paratypes) of all nominal
forms of Notropis, but some lots have doubtless escaped detection, either be-
cause of inadequate labeling or because the series were misplaced on the
shelves (i.e. mixed with nontype material). For inadequately labeled lots,
value judgments must necessarily be made as to whether or not the specimens
should be considered types, this based on agreement of data on the label and/
or elsewhere in the jar with that included in the original description. As might
be anticipated, problems of this sort occur most frequently in the older col-
lections, with the United States National Museum of Natural History being
particularly prominent in this regard.
Listing of all nominal forms is done alphabetically by species or subspecies
name. This arrangement, which is somewhat different from that of Collette
and Knapp (1967), was done to facilitate the finding of species names in the
catalogue. The following data appear under each separate heading: (1) origi-
nal genus and species combination, together with describer and year of de-
scription; (2) abbreviated citation of original description; (3) type locality;
(4) catalogue numbers) of holotype, lectotype, neotype, or syntypes, together
with numbers of specimens, range in standard length in mm, and name of col-
lector and date of collection, if known; (5) catalogue numbers) of paratypes,
number of specimens in each series, and an indication of which are paratopo-
types; (6) remarks concerning the types (counts and condition), nomencla-
tural history of the taxon and definitive references; (7) exact date of original
description if known, or, if not known, the most accurate date determinable;
and (8) present status of the taxon.
In addition to the above list, the following supplementary lists are in-
cluded: (1) a generic synonymy; (2) presently recognized species of Notropis,
with their included subspecies (if any) and junior and senior synonyms (if any);
(3) species presently referred to the genus Hybopsis, but whose affinities are
with Notropis and in which genus they may eventually be placed; (4) nominal
species of Notropis that are based on hybrids, one or both parents of which be-
long to that genus; (5) nominal species of Notropis that are based on hybrids,
BULLETIN FLORIDA STATE MUSEUM
both parents of which belong to other genera; (6) nomina nuda, with their
identifications; and (7) species that were originally described in Notropis, but
which are now referred to other genera. Also, the genus is defined and a brief
discussion of the nomenclatural history of the group is presented, which in-
cludes a comparison of the numbers of forms described during each decade.
The generic synonymy includes 31 names. Each is accompanied by the
describer and date, the type species and its present identification, the method
by which the type species was designated, and the reference source. If the
name is that of a presently recognized subgenus of Notropis, or if it is clearly
a junior synonym of one of these accepted groups, this is also indicated. One
subgenus listed here (Codoma) recently has been elevated tentatively to a full
genus (Miller 1976:12). The status of Opsopoeodus, which Gilbert and Bailey
(1972) downgraded to a subgenus of Notropis, was questioned by Campos and
Hubbs (1973) after they discovered a reduced chromosome number (48 vs 50
pairs) in the species emiliae. Retention here of Codoma and Opsopoeodus as
subgenera of Notropis does not necessarily imply disagreement with Miller
or with Campos and Hubbs. Rather, this course is being followed because
these taxa and their included species have been associated with Notropis so re-
cently, and omitting them from the catalogue would cause the reader incon-
venience and possible confusion. The genus Hybopsis (together with its junior
synonym, Erinemus), which may be included in the synonymy of Notropis in
the future, is listed separately following the generic synonymy of Notropis.
This is to prevent possible misinterpretation that it is being so synonymized
here. Such an action would be premature, inasmuch as the taxonomic rela-
tionships of some species presently placed in Hybopsis have not yet been com-
pletely resolved, and nomenclatural complications would result.
In the list of presently recognized species of Notropis, valid subspecies
are included in subheadings under the species name, together with their re-
spective synonyms. In this list, currently accepted taxonomic decisions (based
on the 1970 American Fisheries Society checklist) are followed in nearly all
cases, except in those situations where recent work clearly indicates that
changes are in order. The taxonomic status of the Mexican forms not included
in the AFS checklist has been determined from consultation with Drs. Robert
R. Miller, University of Michigan, and Salvador Contreras, Universidad Au-
t6noma de Nuevo Leon (see also Contreras 1975; Miller 1976). It has been
necessary in several cases to make arbitrary decisions regarding subspecies,
some of which are not recognized even though they have never been formally
synonymized. Similarly, several taxa are here recognized as full species, de-
spite recent papers in which they are not so treated.
In the above lists, those museums or institutions referred to are indicated
by letter designations as follows: AMNH (American Museum of Natural His-
tory); ANSP (Academy of Natural Sciences of Philadelphia); ASU (Arizona
State University); BCF (United States Bureau of Commercial Fisheries);
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
BMNH (British Museum [Natural History]); BU (Baylor University); CAS or
CAS-SU (California Academy of Sciences and combined California Academy
of Sciences/Stanford University); CU (Cornell University); FMNH (Field
Museum of Natural History [formerly Chicago Natural History Museum]);
FSU (Florida State University); INHS (Illinois Natural History Survey); IPN
(Instituto Politecnico Nacional [Escuela Nacional de Ciencias Biol6gicas de
Mexico]); IU (University of Indiana); KU (University of Kansas); MCZ (Mu-
seum of Comparative Zoology [Harvard University]); MNHN (Museum Na-
tional d'Histoire Naturelle [France]); NCSM (North Carolina State Museum);
NMC (National Museum of Canada [formerly Victoria Memorial Museum]);
OAM (Oklahoma State University [formerly Oklahoma A&M University]);
OSM (Ohio State Museum); TNHC (Texas Natural History Collection [Uni-
versity of Texas]); TU (Tulane University); UAIC (University of Alabama Ich-
thyological Collection); UANL (Universidad Aut6noma de Nuevo Leon [Fac-
ultad de Ciencias Biol6gicas]); UF (University of Florida [Florida State Mu-
seum]); UMMZ (University of Michigan, Museum of Zoology); UNAM (Uni-
versidad Nacional Aut6noma de M6xico [Instituto de Biol6gia]); UOMZ (Uni-
versity of Oklahoma, Museum of Zoology); USFC (United States Fish Com-
mission); and USNM (United States National Museum [now National Museum
of Natural History]).
All original descriptions were consulted to determine whether certain
series comprised type material, inasmuch as syntypes and paratypes occa-
sionally are not so labelled, either on the jars or in the museum catalogue. In
many original descriptions, and occasionally in relatively recent ones, the
number and disposition of syntypes or paratypes is not clearly indicated. Un-
doubtedly the biggest problem in this regard pertains to the types, as well as
the type localities, for the many species Charles Girard described in the late
1850's (118 new cyprinids appeared in his 1857 paper alone). Most of these
fishes were collected during the various United States Government Railroad
and Mexican boundary surveys of a few years earlier, and the conditions
under which the specimens were obtained, together with frequent inade-
quate labeling of the material, have sometimes made it difficult to pinpoint
type localities. (This is discussed further below).
Further compounding the problem is the fact that the number of extant
syntypes of a species is occasionally greater, and frequently less, than the num-
ber listed by Girard (1858, 1859a). I cannot account for the former situation,
but the latter undoubtedly results from the fact that most of Girard's syntypic
series were later broken up into smaller lots and sent on exchange to other
institutions. (This statement, as well as others to follow in this paragraph, are
generally true of most of the older type series initially housed at the USNM).
Those sent to such places as the Academy of Natural Sciences of Philadelphia
and the Museum of Comparative Zoology present no problems, inasmuch as
this material is still extant and readily available. Much material, however,
1978
BULLETIN FLORIDA STATE MUSEUM
cannot be accounted for. Some may have gone to Indiana University, where it
was subsequently destroyed by fire in 1883. Apparently Barton A. Bean, then
Curator of the United States National Museum fish collection, distributed
many USNM types indiscriminantly to various individuals and institutions be-
tween 1892 and 1932. Many of these lots were never returned. Some may also
have been lost at the National Museum because of poor curatorial procedures.
Whatever the reasons, most of Girard's species are represented by fewer type
specimens than were originally present, and in some cases by none at all. Un-
fortunately, those forms for which type material is no longer extant often
cannot be positively identified, as Girard's descriptions are generally quite
vague and could refer to any one of several species.
Type localities are defined as accurately as possible. As might be expected,
there is usually an inverse correlation between age of the species' description
and geographical preciseness of the type locality. Sometimes locality data are
included that did not appear in the original description, and may involve
name changes for existing places or new names for places or political units
that did not exist at the time of description; this is especially true of localities
for species collected during the above-mentioned Railroad and Mexican
boundary surveys. Fortunately, accurate day-by-day accounts were usually
kept for those expeditions, and these, together with maps of the survey routes,
were later published. Reference to these often permits one to pinpoint type
localities for many species, as well as dates of collection. One result of this,
not entirely unexpected, has been the discovery that a number of previously
recognized type localities are erroneous, sometimes so markedly as to warrant
nomenclatural reconsiderations in some cases.
For those species now regarded as junior synonyms, or which have been
so regarded at some time in the past, I have attempted to determine the ref-
erence in which the taxon was first synonymized. These are probably not ac-
curate in every case, as some taxa were synonymized in rather obscure papers
or in papers where this would not normally be expected. Any corrections
noted by the reader are hereby solicited.
I personally have examined over 90 percent of the type lots included in the
present catalogue. This includes the pharyngeal teeth of a number of the
United States National Museum types, which are stored separately in the os-
teological collection, and in some cases are the only part of a type series still
remaining. Those collections with comparatively large holdings that were not
examined include the Field Museum of Natural History, the University of
Kansas, the British Museum (Natural History), and the Museum National
d'Histoire Naturelle (France). Those collections with smaller holdings of types
that were not examined include the American Museum of Natural History,
Natural Museum of Canada, Ohio State Museum, Illinois Natural History
Survey, University of Oklahoma, Oklahoma State University, Texas Natural
History Collection, Arizona State University, Universidad Autonoma de
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
Nuevo Leon, Instituto Politecnico Nacional, and the Universidad Nacional
Autonoma de Mexico.
Information on types in the Field Museum collection appears in Grey
(1947). Additional information on the Field Museum holdings was obtained
from Loren P. Woods and Robert Karl Johnson (in litt). Information on the
Notropis types in the two European museums was obtained through the
courtesy of James Chambers (BMNH) and Marie-Louise Bauchot (MNHN).
William F. Smith-Vaniz, Academy of Natural Sciences of Philadelphia; Royal
D. Suttkus, Tulane University; and Carl L. Hubbs, Scripps Institution of
Oceanography, also generously supplied me with data on BMNH Notropis
types; and Robert E. Jenkins, Virginia Commonwealth University, provided
information on certain Hybopsis types. William I. Follett, California Acad-
emy of Sciences, and C. Richard Robins, University of Miami, have been ex-
tremely helpful in resolving several complicated nomenclatural problems. I
also would like to thank Helen Hauck, Cleveland Public Library, for infor-
mation pertaining to publication of J. P. Kirtland's (1854) paper in the Cleve-
land Annals of Science.
Many individuals have answered questions that have subsequently arisen
regarding types housed at institutions I had visited earlier, and the patience
and cooperation of the following people is gratefully acknowledged: William
Ralph Taylor, James Frank McKinney, and Susan Karnella, National Museum
of Natural History; Myvanwy M. Dick, Museum of Comparative Zoology;
Robert Rush Miller, University of Michigan; Pearl M. Sonoda, California
Academy of Sciences; and William G. Saul and Eugenia B. B6hlke, Academy
of Natural Sciences of Philadelphia. I particularly wish to thank Eugenia
Bohlke, inasmuch as she has called my attention to several situations that,
had they not been noticed, would have resulted in some serious errors in the
catalogue.
Considerable attention has been devoted to verifying publication dates, or,
to be more precise, mailing dates, as this actually determines time of publica-
tion (Mayr, Linsley, and Usinger 1953:222). For most recent papers this pre-
sents no problem, inasmuch as journals now regularly include such informa-
tion, either in each individual number or in the final number of a volume. For
older papers, however, this often is not the case. Fortunately, several publi-
cations exist that provide this sort of information for some of the major mu-
seums (Nolan 1913, Sherborn 1934, True et al. 1946), for the various early
United States government fisheries publications (MacDonald 1921), or for cer-
tain individuals such as Cope (Osborn 1931) and Jordan (Hays 1952). Books
pose a particularly difficult problem, as their exact date of publication is
rarely indicated, even today. When the mailing date for a book or journal
number is not known, the date of publication has been determined according
to Article 21 of the International Code of Zoological Nomenclature (1961),
which says that in the absence of a specified publication date, the earliest
1978
BULLETIN FLORIDA STATE MUSEUM
date of receipt is considered to constitute date of publication (Article 21f);
many of the early publication dates for numbers of the several journals of
the Academy of Natural Sciences of Philadelphia were determined in this way
(Nolan 1913). Some taxa are listed in the present catalogue as having been
described on "(31 December)" of a particular year. According to Article
21b(ii) this date is to be used in all cases in which the day or month of publi-
cation cannot be determined. In cases where the month, but not the day, of
publication is known, the last day of that month is the one automatically desig-
nated [Article 21b(i)], as'for example "(30) November 1931." It has been cus-
tomary for many years for libraries to stamp the date of receipt on or inside
the covers of journals and books as they are received, and it is quite likely that
in some cases earlier description dates will ultimately be found than are indi-
cated here. Hopefully readers will inform me of such cases as they are dis-
covered.
Although space does not permit discussion of every example involving
confused or obscure publication dates, several are of sufficient importance of
interest to merit comment. One of the most complex situations, which in-
volves several of Edward D. Cope's publications appearing in the 1860's, has
been discussed previously (Gilbert 1971a). Although I said nothing in that
paper about the type specimens of those species Cope originally described in
Giinther's (1868) catalogue, questions have since been raised as to (a) whether
only those specimens actually listed in the original description (i.e. those in
the British Museum) should be considered types, or whether the types should
also include the remainder of the series retained by Cope; and (b) whether
the type localities should be limited to those specifically mentioned in the
original description, if part of Cope's material was from other localities. I have
consulted with both William I. Follett and C. Richard Robins on these mat-
ters, and their advice and opinions have been extremely helpful in resolving
the problem. The following facts were considered pertinent in reaching a de-
cision:
(a) the original descriptions obviously were prepared by Cope from all
specimens in his original series, and not merely those sent to the
British Museum;
(b) five species originally described in the catalogue were not represented
by BMNH specimens but were represented by material in Cope's pos-
session;
(c) all specimens clearly were intended to be types, and it was merely
by chance that Giinther's catalogue preceded Cope's papers in pub-
lication;
(d) Girard described 118 new cyprinid species in his 1857 paper, and in
no case were catalogue numbers or the number of type specimens in-
dicated in the original descriptions. He (Girard 1858, 1859a) listed
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
these subsequently, and they have been accepted as the types without
question ever since;
(e) the original description of Hybognathus hankinsoni (Hubbs, in Jordan
1929:88) was not accompanied by either an indication of type speci-
mens or a specific type locality, although the overall range limits of
the species were given. Bailey (1954) regarded as syntypes the speci-
mens Hubbs catalogued earlier as types in the University of Michigan
collection, from which Bailey selected a lectotype and a type locality.
I conclude that there is ample precedent for recognizing all of Cope's orig-
inal specimens as syntypes of those species he described in Gunther's (1868)
catalogue, and not merely those listed in the original accounts. Furthermore,
for those species (i.e. Alburnellus jaculus) in which Cope's original material
came from several localities, but for which individuals from only one locality
were sent to Giinther, all specimens should be regarded as syntypes and all
localities as the potential type locality.
Another complex situation concerns the various U. S. government fisheries
publications appearing in the late 19th and early 20th centuries. These were
often published as house or senate documents or as author's preprints, and the
publication dates of these frequently differ from those appearing in the bound
volume itself. (In such cases the oldest available date is always the one cited.)
Thus Jordan's paper on fishes from the Allegheny region of Virginia, North
Carolina, and Tennessee appeared in the Bulletin of the U. S. Fish Commis-
sion for the year 1888. The date of publication listed for this paper in the
volume is 13 March 1890, and this is the date usually cited, but the article
was issued earlier as a preprint sometime during 1889 (see Hays 1952:116).
As I have been unable to determine the exact day on which the preprint ap-
peared, the publication date should properly be cited as "(31 December)
1889," at least until such time as an earlier date can be verified. Records of
exact publication dates for the early United States National Museum Bulletins
were not kept until after 1906 (True et al. 1946:12-18). The year "1882" ap-
pears on the cover of Bulletin 16, and this is often cited as the year of publi-
cation. However, in the opening paragraph of a later paper by Jordan (1885b:
789) appeared the remark that Bulletin 16 was not actually mailed until 1
April 1883, which thus becomes the official date of publication.
I have chosen not to designate lectotypes in most cases, even where such
designations are clearly indicated, because I think this should be done by in-
dividuals who have conducted a definitive taxonomic study of the species in
question. Several new lectotypes designated in the present catalogue result
from studies that, though completed, are still unpublished (Gibbs 1955 [in
part], Seaman 1968). In such cases, the lectotype designations are credited to
the individuals responsible. Lectotype designations emanating from certain
Fowler (1909, 1910, 1918) papers require special comment because, although
BULLETIN FLORIDA STATE MUSEUM
Fowler has officially been credited with these designations, I am actually the
one responsible for choosing them. In an earlier paper (Gilbert 1971b), I re-
viewed the circumstances surrounding Fowler's lectotypes in my request to
the International Commission on Zoological Nomenclature, in which I asked
that these designations be suspended. The reader is referred to that paper for
pertinent details. Inasmuch as this request will not be approved, for reasons
discussed in my substitute petition (Gilbert 1977), it has become necessary
to select and segregate lectotype specimens from the syntypic series, some-
thing that Fowlder did not do. Fortunately Fowler's illustrated types (i.e. the
lectotypes, according to Article 74b of the International Code), which ap-
peared in his 1910 paper, can be identified in most cases, inasmuch as he char-
acteristically drew the largest specimen in a series and also included a scale of
measurement. Using this scale, the length of the illustrated specimen can be
readily calculated. Fowler's drawings have been found to be surprisingly ac-
curate, so far as their body lengths are concerned, and this, together with the
fact that a number of the series contain one specimen that is clearly longer
than the others, permits accurate selection of Fowler's lectotypes in many
cases. For those series in which no one specimen is notably longer than the
others, it simply becomes a matter of making the best choice possible.
My second request to the International Commission (Gilbert 1977) per-
tains only to Fowler's lectotype designations for two species, Hypsilepis cor-
nutus cerasinus Cope 1868 and Alburnops plumbeolus Cope 1865, both of
which are based on specimens of species other than those to which the orig-
inal descriptions apply and to which the names are applied today. In each
case this resulted from a complex syntypic series, of which the largest speci-
men (and thus the one illustrated) happened to be the "wrong" species.
Fowler's illustration of Hypsilepis cornutus cerasinus (= Notropis cerasinus)
is based on a specimen of Notropis albeolus Jordan 1889, and that of Albur-
nops plumbeolus (= Notropis chrysocephalus [Rafinesque 1820]) on a speci-
men of Notropis heterodon (Cope 1865). Should these lectotype designations
be allowed to stand, the species now called Notropis albeolus would become
N. cerasinus, and the present N. cerasinus would require a new name. The
consequences- of retaining the specimen of Notropis heterodon as lectotype of
Alburnops plumbeolus are less critical, although it is conceivable that the
long-established name Notropis heterodon could be changed to N. plumbeolus
should the first reviser show imprudent judgment.
During this study, several cases of incorrect synonymization were dis-
covered, although only two involve potential name changes for recognized
species. These are: Cyprinella ludibunda Girard 1857, a senior synonym of
Notropis stramineus (Cope 1865) and Notropis volucellus (Cope 1865) (the
two extant syntypes comprise one specimen of each species); and Notropis
phenacobius Forbes 1885, a probable senior synonym of Notropis amnis
Hubbs and Greene 1951 (identification based on text of original description
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
and accompanying figure, geographic locality, and other circumstantial evi-
dence). Inasmuch as other ichthyologists are currently studying the species
involved, I have elected to take no further action on these matters.
DEFINITION OF GENU'S Notropis
The genus Notropis Rafinesque (1818) is a large and diverse group of Cy-
prinidae containing numerous closely related, morphologically similar forms.
At various times in the past its principal components have been regarded as
distinct, though poorly defined genera. Gilbert (1884a) was the first to pro-
pose that a number of these be combined under the name Notropis. Jordan
(1885b) relegated these groups to subgeneric status within the broad genus
Notropis, where they (together with a number of subgenera subsequently pro-
posed) have remained since, despite sporadic efforts (Jordan 1929, Jordan,
Evermann, and Clark 1930) to separate them. Although numerous new spe-
cies have been described down through the years, only one (Notropis emiliae)
has so far been added through downgrading of a genus (Opsopoeodus) to a sub-
genus (Gilbert and Bailey 1972). A second species harperri, which was long
associated with the genus Hybopsis, was recently reassigned to Notropis (Gil-
bert and Bailey 1972), the genus in which it was originally described. About
eight other barbelled species, currently also included in Hybopsis, may even-
tually be removed and incorporated into the group. One subgenus (Codoma),
comprising a single species, is in the process of being elevated to full generic
status (R. R. Miller, pers. comm.), and the allocation of Opsopoeodus to No-
tropis was recently questioned by Campos and Hubbs (1973). It has also been
suggested that the subgenus Cyprinella, comprising 19 species, may be suf-
ficiently distinct to merit generic recognition (R. E. Jenkins, pers. comm.). Of
those genera presently regarded as distinct and which are not treated in this
catalogue, Dionda (with eight species; Hubbs and Miller 1974, 1977, Con-
treras and Verduzco 1977) is the most likely candidate for eventual inclusion
in Notropis.
Until recently the genus Notropis could be characterized by the following
characters: (a) always one or two rows of pharyngeal teeth, and never with
more than four in the greater (major) row; (b) dorsal-fin rays normally eight
in number; (c) gut always short; (d) no maxillary barbel present at corners of
mouth; (e) scales of relatively large size, never numbering more than 55 in the
lateral series and usually fewer than 40; and (f) body size generally small, usu-
ally less than 100 mm standard length and never exceeding 175 mm SL.
Several papers have appeared in recent years that have caused the former
cohesiveness of the group to be broken down. Cort6s (1968), in her descrip-
tion of Notropis imeldae, noted that the species usually has a barbel at each
corner of the mouth. Jenkins and Lachner (1971) and Gilbert and Bailey
(1972) discussed the artificiality of this character, pointing out that barbels
BULLETIN FLORIDA STATE MUSEUM
sometimes may be present or absent within a single species, that they have
undergone parallel evolution in many diverse cyprinid lines (presumably in
response to certain environmental stimuli), and that although they usually
are diagnostic of individual species, they are of limited value in the delinea-
tion of genera.
Swift (1970, 1971) recommended inclusion of Dionda nubila in Notropis,
noting that this species bears close resemblance to the subgenus Hydrophlox
except for having a long, coiled gut and herbivorous feeding habits. Snelson
(1971) demonstrated a similar condition for Notropis mekistocholas, a species
whose closest relatives each has a short gut.
Gilbert and Bailey (1972) downgraded Opsopoeodus (with one species and
two subspecies) to a subgenus of Notropis, and in so doing eliminated two
other characters by which the genus had formerly been distinguished. No-
tropis emiliae characteristically has nine dorsal rays, as opposed to eight in
all other members of the genus presently described. The reduced importance
of this character as a diagnostic feature of Notropis is further substantiated by
the usual presence of 9 or 10 dorsal rays in an undescribed species that is ob-
viously closely related to N. welaka. N. emiliae has always been characterized
as having five pharyngeal teeth in the greater row of each pharyngeal arch,
but Gilbert and Bailey (1972) showed their distinctive new subspecies, N.
emiliae peninsularis, to have a count of four in the right arch. In addition,
they presented morphological evidence to substantiate their idea that the
upper (fifth) tooth in N. emiliae was derived from an ancestral condition of
four, thus reversing an earlier phyletic reduction.
In summary, the genus Notropis can no longer be so readily characterized
as in the past. Of the six diagnostic characters listed above, only those relating
to body and scale size have not been modified or refuted in recent years. Al-
though it is true that no species of Notropis has more than two rows of pharyn-
geal teeth, the same can be said for all native North American Cyprinidae. On
the other hand, it should also be pointed out that the presumed addition of a
tooth in the greater row in N. emiliae means that the original diagnostic
pharyngeal-tooth character (i.e. four teeth in this row) does, in essence, still
hold. Possibly investigations now under way will reveal other characters that
will better unify the genus, or, conversely, will serve to dismember it. Among
these are nuptial tuberculation, patterns of gut coiling, serum protein anal-
yses, karyology, and osteology.
TAXONOMIC HISTORY OF GENUS Notropis
The taxonomic history of the genus Notropis parallels in large degree that
of the darters (family Percidae), which was earlier reviewed by Collette
(1967), and also reflects the overall history of North American Ichthyology
during the past 160 years. Collette divided activity in darter systematics into
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
four periods ("Beginnings" [1818-1840]; "Golden Era" [1841-1892]; "Dol-
drums" [1893-1931]; and "Reawakening" [1932-present]), and these are (with
certain modifications in dates) equally appropriate here. A comparison of the
number of new Notropis forms (both valid and invalid) appearing during each
decade is shown in Table 1.
The first species of Notropis to be described was Cyprinus cornutus
Mitchill 1817. This was followed by 13 others during the next seven years
(Rafinesque 1817, 1818, 1820a, 1820b; Clinton 1824), of which only three (No-
tropis atherinoides Rafinesque 1818, Luxilus chrysocephalus Rafinesque 1820,
and Clupea hudsonia Clinton 1824) are regarded as valid today. The primary
reason for the plethora of invalid forms is the inadequacy of Rafinesque's
descriptions, and this, coupled with a total lack of type material, makes most
of his taxa impossible to identify. Although some of Rafinesque's original notes
and drawings still exist and are of value in identifying some of the more dis-
tinctive fish species (see Collette and Knapp 1967: Figs. 1, 3, and 5), they are
too crude in most cases to permit recognition of the various kinds of Cyprini-
dae.
TABLE 1.-SUMMABY BY DECADE OF NUMBERS OF SPECIFIC AND SUBSPECIFIC NAMES PHO-
POSEI) THAT ARE NOW REFERABLE TO THE (GENS lV)tropiS.
Decade Valid Invalid Total
beginning names names names
*1810 2 2 4
1820 2 8 10
1830 0 0 0
1840 0 6 6
1850 16 38 54
1860 23 19 42
1870 23 18 41
1880 14 24 38
1890 9 17 26
1900 4 9 13
1910 0 3 3
1920 7 3 10
1930 1 0 1
1940 5 12 17
1950 13 5 18
1960 5 0 5
**1970 7 0
TOTALS 131 164 295
*First name proposed in 1817.
'New names proposed as of September 1977.
1978
BULLETIN FLORIDA STATE MUSEUM
Following this initial flurry of activity, no other new species of Notropis
appeared until 1842, at which time Leuciscus vittatus DeKay 1842 (a junior
synonym of Notropis cornutus) was described. This was followed by Argyreus
rubripinnis Heckel 1843 and Leuciscus spirlingulus Valenciennes 1844, both
of which are also junior synonyms of N. cornutus. Storer (1845) published de-
scriptions of other new forms, at least three of which likely belong to Notropis
but none of which can positively be identified to species.
The 1850's showed a sharply increased level of activity (the beginning of
the Golden Era for the genus Notropis). This was initiated by two lengthy
publications by Agassiz (1850, 1854), in which he described several new taxa,
and by a series of short descriptions by Baird and Girard (1853), Kirtland
(1854), and Lapham (1854). These were followed by a veritable deluge of new
Notropis species a few years later, 44 of which first appeared in Girard's 1857
paper. Despite the large number of new species described during this decade,
comparatively few are considered to be valid today (16 of 54, or 29.6%). This
stems largely from the highly uncritical nature of Girard's descriptions: 33 of
the 44 new forms described in his 1857 paper are not considered valid, and
14 are junior synonyms of Notropis lutrensis.
Descriptions of new Notropis species continued unabated over the next
40 odd years, beginning with Edward D. Cope's eight publications between
1865 and 1875, continuing with a multitude of papers (mostly by David S.
Jordan and students) from 1876 through 1899, and culminating with several
papers, dealing primarily with new Mexican species, between 1902 and 1904.
The end of the Golden Era actually was signaled a few years earlier by the
publication of Jordan and Evermann's (1896-1900) monumental four volumes
on the "Fishes of North and Middle America." This work, although of tre-
mendous importance to Ichthyology, resulted in the premature decline of
systematic work on North American freshwater fishes, inasmuch as Jordan
regarded studies in this area to be essentially complete. Very few new species
were described during the next 20 years (the Doldrum period), only seven
new forms (two valid) appearing from 1905 through 1925. The mid-1920's sig-
nify the period of Reawakening. This largely resulted from the activities of
Carl L. Hubbs who, with his associates, described six of the eight new forms
(five valid) that appeared from 1926 through 1929.
The period of Reawakening is considered to have begun in 1926, but if
one ignores the 4-year flurry of activity in the late 1920's, the Doldrum period
actually may be regarded as lasting through the 1930's so far as Notropis sys-
tematics (but not North American freshwater ichthyology as a whole) are con-
cerned. Only one new taxon (Notropis volucellus wickliffi Trautman) ap-
peared during the 1930's, the second lowest total for any decade of the 17
under consideration (which cover 160 years). The 1940's saw a sharply in-
creased degree of activity in the study of North American cyprinid fishes in
general and the genus Notropis in particular, which has continued unabated
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
to the present day. The relatively small number of new forms described dur-
ing the 1960's and 1970's, as compared to the 1940's and 1950's, may be ac-
counted for both by the present state of knowledge (fewer new species are
now being discovered), as well as a slightly more conservative philosophy re-
garding the description of new subspecies. At present, between 12 and 15
species of Notropis are known that remain to be described (Jenkins 1976), and
subsequent study probably will reveal several more that are present in col-
lections but which have not yet been distinguished.
GENERIC SYNONYMY'
Notropis Rafinesque 1818:204 (type species Notropis atherinoides Rafinesque 1818, by mono-
typy). (= Notropis)
Minnilus Rafinesque 1820b:45 (type species Minnilus dinemus Rafinesque 1820b [= Notropis
atherinoides Rafinesque 1818], by subsequent designation of Jordan and Gilbert 1877:86).
(= Notropis)
Luxilus Rafinesque 1820b:47 (type species Luxilus chrysocephalus Rafinesque 1820b, by sub-
sequent designation of Jordan and Gilbert 1877:86). (= Luxilus)
Plargyrus Rafinesque 1820b:50-51 (type species Rutilus plargyrus Rafinesque 1820b [= Luxilus
chrysocephalus Rafinesque 1820b], by absolute autonomyy. (= Luxilus)
Hypsolepis Baird, in Agassiz 1854:359 (type species Cyprinus cornutus Mitchill 1817, by original
designation; spelling emended by Cope [1865a:279] to Hypsilepis). (= Luxilus)
Alburnellus Girard 1857:193 (type species Alburnus dilectus Girard 1857 [ = Notropis atherinoides
Rafinesque 1818], by subsequent designation of Jordan and Gilbert 1877:91). (= Notropis)
Alburnops Girard 1857:194 (type species Alburnops blennius Girard 1857, by subsequent desig-
nation of Jordan and Gilbert 1877:91). (= Alburnops)
Codoma Girard 1857:194-195 (type species Codonma ornata Girard 1857, by subsequent desig-
nation of Jordan and Gilbert 1877:91; tentatively recognized by Miller [1976] as a valid
genus). (= Codonma)
Cyprinella Girard 1857:196-197 (type species Leuciscus bubalinus Baird and Girard 1853
[= Leuciscus lutrensis Baird and Girard 1853], by subsequent designation of Jordan and
Gilbert 1877:91). (= Cyprinella)
Moniana Girard 1857:199 (type species Leuciscus lutrensis Baird and Girard 1853, by sub-
sequent designation of Jordan and Gilbert 1877:91). (= Cyprinella)
Hudsonius Girard 1857:210 (type species Clupea hudsonia Clinton 1824, by subsequent desig-
nation of Jordan and Gilbert 1877:92).
Graodus Giinther 1868:485-486 (type species Graodus nigrotaeniatus Giinther 1868 [= Leuciscus
boucardi Giinther 1868], by monotypy).
Photogenis Cope 1868a:163-164 (type species Leuciscus spilopterus Cope, in Giinther 1868, by
subsequent designation of Cope, in Jordan and Copeland 1877:154). (= Cyprinella)
Lythrurus Jordan 1876b:271-272 (type species Semotilus diplemiia Rafinesque 1820b, which is
actually Hypsilepis diplaemia Cope 1868a [=Lythrurus cyanocephalus Copeland, in
Jordan 1877a], by subsequent designation of Jordan and Gilbert 1877:95). (= Lythrurus)
Episema Cope and Jordan, in Jordan 1877b:77-78 (type species Photogenis scabriceps Cope 1868a,
by original designation; preoccupied by Episema Ochsenheimer 1816, a genus of lepi-
dopteran insect).
'Includes only those nominal genera presentlv placed in synonymy of otropis. Those genera that may ultimately he synon-
ymized with Notropis are listed separately following this account.
BULLETIN FLORIDA STATE MUSEUM
Nototropis Jordan 1877c:342-343 (type species Nototropis stilbius Jordan 1877c, herein desig-
nated). (= Notropis)
Chriope Jordan 1878c:787 (type species Hybopsis bifrenatus Cope 1869, by original designa-
tion).
Hydrophlox Jordan, in Jordan and Brayton 1878:18-19 (type species Hybopsis rubricroceus Cope
1868b, by original designation). (= Hydrophlox)
Erogala Jordan, in Jordan and Brayton 1878:20-21 (type species Photogenis stigmaturus Jordan
1877c[ = Cyprinella venusta Girard 1857], by original designation). (= Cyprinella)
Opsopoeodus Hay 1881:507 (type species Opsopoeodus emiliae Hay 1881, by monotypy; recog-
nized by Campos and Hubbs [1973] as a valid genus). (= Opsopoeodus)
Coccotis Jordan 1882:852 (type species Leuciscus coccogenis Cope, in Gunther 1868, by original
designation). (= Luxilus)
Trycherodon Forbes, in Jordan and Gilbert 1883:247 (type species Trycherodon megalops
Forbes, in Jordan and Gilbert 1883 [= Opsopoeodus eniliae Hay 1881], by original desig-
nation). (= Opsopoeodus)
Miniellus Jordan 1888:56 (type species Hybognathus procne Cope 1865a, by subsequent desig-
nation of Jordan 1920:442).
Opsopoea Jordan and Evermann 1896:249 (type species Opsopoeodus bollhani Gilbert 1890
[ = Hemitremia maculata Hay 1881], by original designation).
Azteca Jordan and Evermann 1896:254, 258 (type species Codoma vittata Girard 1857 [= Cera-
tichthys sallaei Giinther 1868], by original designation; Codomna vittata Girard 1857 is pre-
occupied in Notropis by Leuciscus vittatus DeKay 1842). (= Aztecula)
Orcella Jordan and Evermann 1896:254, 289 (type species Notropis orca Woolman 1894 [= Albur-
nellus simus Cope, in Cope and Yarrow 1875], by original designation).
Aztecula Jordan and Evermann 1898:2799 (type species Codoma vittata Girard 1857 [= Cera-
tichthys sallaei Ginther 1868]; substitute name for Azteca, Jordan and Evermann 1896, pre-
occupied by Azteca Forel 1878, a genus of hymenopteran insect). (= Aztecula)
Orcula Jordan and Evermann 1900:3410 (type species Notropis orca Woolman 1894 [=Albur-
nellus simus Cope, in Cope and Yarrow 1875]; substitute name for Orcella, Jordan and Ever-
mann 1896, incorrectly regarded as preoccupied by Orcaella Gray 1866, a genus of ce-
tacean mammal). (= Orcella)
Paranotropis Fowler 1904b:245 (type species Photogenis leuciodus Cope 1868a, by original desig-
nation). (= Hydrophlox)
Coccogenia Cockerell and Callaway 1909:190-191 (type species Leuciscus coccogenis Cope, in
Ginther 1868, by original designation). (= Luxilus)
Pteronotropis Fowler 1941:234 (type species Leuciscus hypselopterus Ginther 1868, by mono-
typy; proposed as a subgenus of Notropis). (= Pteronotropis)
STATUS OF GENUS Nototropis
Some confusion attends the nominal genus Nototropis. The genus was first
proposed by Jordan (1877c:342-343), who included two species (N. lirus and
N. stilbius), both of which were described as new but neither of which was
specifically designated as the type species. Although consistent spelling of the
name throughout the paper eliminates the possibility that it was based on a
misspelling of Notropis, it is curious that Jordan (1888, 1920, and other papers)
and Jordan and Evermann (1896) made no subsequent reference to it. Meek
(1904:62-63) later resurrected Nototropis, without comment, including in it
all Mexican species now referred to Notropis. Ruthven (1906) apparently was
the only other author to use the name.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
As no type species is yet available for Nototropis, I hereby so designate
Notropis stilbius, which is a close relative of Notropis atherinoides Rafinesque
1818, the type species of Notropis. Thus Nototropis becomes a junior synonym
of that taxon, both as a genus and subgenus.
GENERA THAT MAY ULTIMATELY BE INCLUDED IN Notropis
Hybopsis Agassiz 1854:358 (type species Hybopsis gracilis Agassiz 1854 [=Rutilus amblops
Rafinesque 1820b], by subsequent designation of Jordan and Gilbert 1877:90). (= Hybopsis)
Erinemus Jordan 1876b:279 (type species Ceratichthys hyalinus Cope, in Giinther 1868:179-
180 [=Rutilus amblops Rafinesque 1820b], by subsequent designation of Jordan and
Gilbert 1877:95). (= Hybopsis)
TAXONOMIC LISTS
VALID FORMS OF Notropis, INCLUDING SYNONYMS' 2
1. Notropis aquirrepequenoi Contreras and Rivera 1973-none.
2. Notropis albeolus Jordan 1889-none.
3. Notropis alborus Hubbs and Raney 1947-none.
4. Notropis altipinnis (Cope 1870)-Notropis altipinnis chowanus Hubbs and Raney 1948;
Notropis altipinnis neusensis Hubbs and Raney 1948; Notropis altipinnis tarensis
Hubbs and Raney 1948; Notropis altipinnis white Hubbs and Raney 1948; No-
tropis altipinnis wright Hubbs and Raney 1948.
5. Notropis atmabilis (Girard 1857)-Alburnus inegalops Girard 1857; Alburnus socius Girard
1857; Cyprinella luxiloides Girard 1857 (?); Cyprinella macrostoma Girard 1857;
Notropis swaini Jordan and Gilbert 1885.
6. Notropis amnis Hubbs and Greene 1951-Notropis phenacobius Forbes 1885 (a probable
senior synonym); Notropis nocomis Evermann 1892 (a preoccupied senior synonym);
Notropis amnis pinnosa Hubbs and Bonham 1951.
7. Notropis amoenus (Abbott 1874)-none.
8. Notropis analostanus (Girard 1859)-none.
9. Notropis anogenus Forbes 1885-none.
10. Notropis gardens (Cope 1868)-Rutilus ruber Rafinesque 1820 (?); Alburnellus matutinus
Cope 1870; Notropis umbratilisfasciolaris Gilbert 1891.
11. Notropis arionmmus (Cope 1868)-none.
12. Notropis asperifrons Suttkus and Raney 1955-none.
13. Notropis atherinoides Rafinesque 1818-Minnilus dinemus Rafinesque 1820; Alburnus
nitidus Kirtland 1854; Alburnus acutus Lapham 1854; Alburnus dilectus Girard
1857; Alburnus lepidulus Girard 1857 (?); Alburnus oligaspis Cope 1865; Notropis
atherinoides caddonis Meek 1891; Notropis louisianae Evermann 1898.
14. Notropis atrapiculus Snelson 1972-none.
15. Notropis atrocaudalis Evermann 1892-none.
16. Notropis bailey Suttkus and Raney 1955-none.
Nominal forms are presumed to be junior synonyms unless stated otherwise.
The systematics of the Notropis lutrensis complex have been clarified only in the southern part of the range (Mexico and southern
Texas) (Contreras 1975). Consequently, the nominal forms described from more northerly areas cannot presently be assigned
to subspecies, and thus are simply listed here as junior synonyms of Notropis lutrensis.
SFossil species.
The systematics of Notrpnis stramineus have been clarified only in one area (South Dakota) (Bailey and Allum 1962). Con-
sequently, only those nominal forms whose subspecific allocation was clearly indicated by those authors are listed by sub-
species here.
1978
BULLETIN FLORIDA STATE MUSEUM
17. Notropis bairdi Hubbs and Ortenburger 1929-none.
18. Notropis bellus (Hay 1881)
a. Notropis bellus alegnotus Snelson 1972-none.
b. Notropis bellus bellus (Hay 1881)-Notropis alabamae Jordan and Meek 1884.
19. Notropis bifrenatus (Cope 1869)-Notropis cayuga Meek 1889.
20. Notropis blennius (Girard 1857)-Episema jejuna Forbes 1878; Notropis albeolus Eigen-
mann and Eigenmann 1893; Notropis jordanii Eigenmann and Eigenmann 1893.
21. Notropis boops Gilbert 1884-none.
22. Notropis boucardi (Giinther 1868)-Ceratichthys cumingii Giinther 1868; Graodus nigro-
taeniatus Ginther 1868.
23. Notropis braytoni Jordan and Evermann 1896-Moniana nitida Girard 1857 (a preoccu-
pied senior synonym); Notropis robustus Meek 1902.
24. Notropis buccula Cross 1953-none.
25. Notropis buchanani Meek 1896-none.
26. Notropis caeruleus (Jordan 1877)-none.
27. Notropis calientis Jordan and Snyder 1899-none.
28. Notropis callisema (Jordan 1877)-none.
29. Notropis callistius (Jordan 1877)-none.
30. Notropis callitaenia Bailey and Gibbs 1956-none.
31. Notropis camurus (Jordan and Meek 1884)-none.
32. Notropis cerasinus (Cope 1868)-none.
33. Notropis chalybaeus (Cope 1869)-Notropis chalybaeus abbotti Fowler 1904.
34. Notropis chihuahua Woolman 1892-none.
35. Notropis chiliticus (Cope 1870)-none.
36. Notropis chloristius (Jordan and Brayton 1878)-none.
37. Notropis chlorocephalus (Cope 1870)-none.
38. Notropis chrosomus (Jordan 1877)-none.
39. Notropis chrysocephalus (Rafinesque 1820)
a. Notropis chrysocephalus chrysocephalus (Rafinesque 1820)-Rutilus plargyrus Rafinesque
1820 (?); Leuciscus obesus Storer 1845 (?); Plargyrus typicus Girard 1857(?); Alburnops
plumbeolus Cope 1865; Hypsilepis corutus gibbus Cope 1868 (possibly based on a hy-
brid: Notropis chrysocephalus X Notropis cornutus); Hybopsis lacertosus Cope 1868.
b. Notropis chrysocephalus isolepis Hubbs and Brown 1927-Leuciscus gibbosus Storer 1845
(?)(a possible senior synonym).
40. Notropis coccogenis (Cope 1868)-Notropis brimleyi Bean 1903.
41. Notropis cornutus (Mitchill 1817)-Cyprinus megalops Rafinesque 1817; Cyprinus mela-
nurus Rafinesque 1817; Cyprinus haematopterus Rafinesque 1820; Cyprinus trivit-
tatus Rafinesque 1820; Rutilus compressus Rafinesque 1820 (?); Leuciscus vittatus
DeKay 1842; Argyreus rubripinnis Heckel 1843; Leuciscus spirlingulus Valenciennes
1844; Leuciscus frontalis Agassiz 1850; Leuciscus gracilis Agassiz 1850; Plargyrus
bowmani Girard 1857; Plargyrus argentatus Girard 1857; Hypsilepis cornutus cya-
neus Cope 1868; Notropis universitatis Evermann and Cockerell 1909.
42. Notropis cummingsae Myers 1925-Notropis cummingsae collis Hubbs and Raney 1951.
43. Notropis dorsalis (Agassiz 1854)-Photogenis piptolepis Cope 1871; Hybopsis montanus
Meek 1885; Notropis gilbert Jordan and Meek 1885; Notropis keimi Fowler 1909;
Notropis horatii Cockerell 1911.
44. Notropis edwardraneyi Suttkus and Clemmer 1968-none.
45. Notropis emiliae (Hay 1881)
a. Notropis emiliae emiliae (Hay 1881)-Trycherodon megalops Forbes 1883; Opsopoeodus
osculus Evermann 1892.
b. Notropis emiliae peninsularis Gilbert and Bailey 1972-none.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
46. Notropis euryzonus Suttkus 1955-none.
47. Notropis fumreus Evermann 1892-Notropis macrolepidotus Forbes 1885 (?) (a possible
senior synonym).
48. Notropis galacturus (Cope 1868)-none.
49. Notropis gibbsi Howell and Williams 1971-none.
50. Notropis girardi Hubbs and Ortenburger 1929-none.
51. Notropis greenei Hubbs and Ortenburger 1929-none.
52. Notropis harper Fowler 1941-Notropis norrisi Fowler 1945; Hybopsis harper subterranea
Hubbs and Crowe 1956.
53. Notropis heterodon (Cope 1865)-none.
54. Notropis heterolepis Eigenmann and Eigenmann 1893-Notropis muskoka Meek 1899; No-
tropis kendalli Evermann and Cockerell 1909; Notropis heterolepis regalis Hubbs
and Lagler 1949.
55. Notropis hudsonius (Clinton 1824)-Hudsonius amarus Girard 1857; Hudsonius fluviatilis
Girard 1857; Hybopsis phaenna Cope 1865; Luxilus selene Jordan 1877; Alburnops
saludanus Jordan and Brayton 1878; Hudsonius euryopa Bean 1880; Notropis sco-
piferus Eigenmann and Eigenmann 1893; Opsopoeodus borealis Harper and
Nichols 1919 (possibly based on hybrid involving Notropis hudsonius).
56. Notropis hypselopterus (Ginther 1868)
a. Notropis hypselopterus hypselopterus (Ginther 1868)-Alburnus formosus Putnam
1863 (a preoccupied senior synonym); Photogenis grandipinnis Jordan 1877; No-
tropis metallicus Jordan and Meek 1884.
b. Notropis hypselopterus stonei Fowler 1921-none.
57. Notropis hypsilepis Suttkus and Raney 1955-none.
58. Notropis imeldae Cort6s 1968-none.
59. Notropis jemezanus (Cope 1875)-Notropis santarosaliae Meek 1902.
60. Notropis leedsi Fowler 1942-none.
61. Notropis lepidus (Girard 1857)-none.
62. Notropis leuciodus (Cope 1868)-none.
63. Notropis lirs (Jordan 1877)-none.
64. Notropis longirostris (Hay 1881)-Leuciscus croceus Storer 1845 (?) (a possible senior syn-
onym).
65. Notropis lutipinnis (Jordan and Brayton 1878)-none.
`66. Notropis lutrensis (Baird and Girard 1853)-Leuciscus bubalinus Baird and Girard 1853;
Cyprinella beckwithi Girard 1857; Cyprinella gunnisoni Girard 1857; Cyprinella
suavis Girard 1857; Cyprinella umbrosa Girard 1857; Moniana leonina Girard 1857;
Moniana frigid Girard 1857; Moniana pulchella Girard 1857; Cyprinella billing-
siana Cope 1871; Moniana jugalis Cope 1871; Cyprinella forbesi Jordan 1878; No-
tropis lutrensis blairi Hubbs 1940.
a. Notropis lutrensis forlonensis (Meek 1904)-none.
b. Notropis lutrensis formosus (Girard 1857)-Notropis mearnsi Snyder 1915.
c. Notropis lutrensis garmani Jordan 1885-Cyprinella rubripinna Garman 1881 (a pre-
occupied senior synonym).
d. Notropis lutrensis lutrensis (Baird and Girard 1853)-Moniana complanata Girard 1857;
Moniana laetabilis Girard 1857; Moniana couchi Girard 1857; Moniana gibbosa Girard
1857; Moniana gracilis Girard 1857; Hypsilepis iris Cope 1875; Cliola montiregis Cope
1885.
e. Notropis lutrensis santamariae Evermann and Goldsborough 1902-none.
67. Notropis maculatus (Hay 1881)-Opsopoeodus bollmani Gilbert 1890; Notropis louisae
Fowler 1940; Notropis burchi Fowler 1942.
'68. Notropis megalepis Smith 1962-none.
1978
BULLETIN FLORIDA STATE MUSEUM
69. Notropis mekistocholas Snelson 1971-none.
70. Notropis moralesi DeBuen 1956-none.
71. Notropis nazas (Meek 1904)-none.
72. Notropis niveus (Cope 1870)-none.
73. Notropis nubilus (Forbes 1878)-none.
74. Notropis ornatus (Girard 1857)-none.
75. Notropis ortenburgeri Hubbs 1927-none.
76. Notropis oxyrhynchus Hubbs and Bonham 1951-none.
77. Notropis ozarcanus Meek 1891-none.
78. Notropis perpallidus Hubbs and Black 1940-none.
79. Notropis petersoni Fowler 1942-Notropis waccamanus Fowler 1942; Notropis williami
Fowler 1945.
80. Notropis photogenis (Cope 1865)-Photogenis leucops Cope 1868; Photogenis leucops en-
graulinus Cope 1868; Alburnellus large Cope 1869.
81. Notropis pilsbryi Fowler 1904-none.
82. Notropis potter Hubbs and Bonham 1951-none.
83. Notropis procne (Cope 1865)-Hybopsis longiceps Cope 1868.
84. Notropis proserpinus (Girard 1857)-Moniana aurata Girard 1857.
85. Notropis pyrrhomelas (Cope 1870)-none.
86. Notropis roseipinnis Hay 1885-Minnilus rubripinnis Hay 1881 (a preoccupied senior
synonym).
87. Notropis rubellus (Agassiz 1850)-Cyprinella lugubris Girard 1857 (?); Alburnus rubrifrons
Cope 1865; Leuciscus copii Giinther 1868; Alburnellus jaculus Cope 1868; Albur-
nellus micropteryx Cope 1868; Alburnellus percobromus Cope 1871.
88. Notropis rubricroceus (Cope 1868)-none.
89. Notropis rutilus Girard 1857-none.
90. Notropis sabinae Jordan and Gilbert 1886-none.
91. Notropis saladonis Hubbs and Hubbs 1958-none.
92. Notropis sallei (Giinther 1868)-Codomna vittata Girard 1857 (a preoccupied senior syn-
onym); Notropis aztecus Woolman 1894; Notropis lermae Evermann and Golds-
borough 1902; Aztecula mexicana Meek 1902.
93. Notropis scabriceps (Cope 1868)-none.
94. Notropis scepticus (Jordan and Gilbert 1883)-none.
95. Notropis semperasper Gilbert 1961-none.
96. Notropis shumardi (Girard 1857)-Alburops illecebrosus Girard 1857; Notropis chamber-
laini Evermann 1898; Notropis hrazosensis Hubbs and Bonham 1951.
97. Notropis signipinnis Bailey and Suttkus 1952-none.
98. Notropis simus (Cope 1875)-Notropis orca Woolman 1894.
99. Notropis spectrunculus (Cope 1868)-none.
100. Notropis spilopterus (Cope 1868)-Hybopsis fretensis Cope 1869; Notropis spilopterus hyp-
sisomatus Gibbs 1958.
101. Notropis stilbius (Jordan 1877)-none.
'102. Notropis stramineus (Cope 1865)-Minnilus microstonmus Rafinesque 1820 (?) (a possible
senior synonym); Cyprinella ludibunda Girard 1857 (a senior synonym of either
Notropis stramineus or Notropis volucellus, as determined from future designation
of lectotype); Alburnus lineolatus Agassiz 1863 (a probable senior synonym); No-
tropis reticulatus Eigenmann and Eigenmann 1893.
a. Notropis stramineus missuriensis (Cope 1871)-Hybopsis scylla Cope 1871; Cliola chlhor
Jordan 1878.
b. Notropis stramineus stramineus (Cope 1865)-(most or all the remaining forms listed
above under the synonymy of Notropis stramineus may ultimately be shown to be-
long to this subspecies).
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
103. Notropis telescopes (Cope 1868)- Notropis telescopes arransanus Meek 1891.
104. Notropis texanus (Girard 1857)-Moniana deliciosa Girard 1857; Luxilus roses Jordan
1877; Notropis nux Evermann 1892; Notropis heterodon richardsoni Hubbs and
Greene 1926; Hudsonius aletes Jordan and Evermann 1927.
105. Notropis topeka (Gilbert 1884)-Notropis aeneolus Hay 1887.
106. Notropis trichroistius (Jordan and Gilbert 1878)-none.
107. Notropis tropics Huhbs and Miller 1975-none.
108. Notropis umbratilis (Girard 1857)
a. Notropis mhbratilis cyanocephalus (Copeland 1877)-Hypsilepis diplaemia Cope 1868
(a senior synonym); Lythrurus atripes Jordan 1878; Minnilus punctulatus Hay 1881;
Notropis lythrunts Jordan 1884; Notropis inacrolepidotus Forbes 1885 (?); Notropis
notemigonoides Evermann 1892.
b. Notropis umbratilis umhratilis (Girard 1857)-Luxilus lucidus Girard 1857; Minnilus
nigripinnis Gilbert 1884.
109. Notropis uranoscopus Suttkus 1959-none.
110. Notropis renustus (Girard 1857)
a. Notropis renustus cercostigma (Cope 1868)-Photogenis eurystomus Jordan 1877; Pho-
togenis leucopus Jordan and Brayton 1878; Luxilus chickasatensis Hay 1881.
bI. Notropis renustus stigmaturus (Jordan 1877)-Cyprinella calliura Jordan 1877 (based
on intergrades: N. t. rercostigmua N. t. stigmaturus).
c. Notropis cenustus renustus (Girard 1857)-Cyprinella notata Girard 1857; Cliola uro-
stigma Jordan and Meek 1884; Notropis cooglei Hildebrand and Towers 1928.
111. Notropis volucellus (Cope 1865)-Minnilus microstomus Rafinesque 1820 (?) (a possible
senior synonym); Cyprinella ludibunda Girard 1857 (a senior synonym of either No-
tropis stramineus or Notropis rolucellus, as determined from future designation of lec-
totype); Notropis nocomis Jordan and Gilbert 1886.
112. Notropis welaka Evermann and Kendall 1898-none.
113. Notropis whipplei (Girard 1857)-none.
114. Notropis wickliffi Trautman 1931-none.
115. Notropis xaenocephalus (Jordan 1877)-none.
116. Notropis xaenurus (Jordan 1877)-none.
117. Notropis xanthicara Minckley and Lytle 1969-none.
118. Notropis zonatus (Agassiz 1863)-none.
119. Notropis mnistius (Jordan 1880)-none.
Valid forms of Hybopsis, including synonyms,
that may eventually be placed in Notropis
1. Hybopsis ambhops (Rafinesque 1820)-Hybopsis gracilis Agassiz 1854; Ceratichthys hya-
linus Cope 1868.
2. Hyhopsis hypsinotus (Cope 1870)-none.
3. Hybopsis labrosa (Cope 1870)-none.
4. Hybopsis lineapunctata Clemmer and Suttkus 1971-none.
5. Hybopsis rubrifrons (Jordan 1877)-none.
6. Hybopsis winchelli Girard 1857-none.
7. Hybopsis zanenma (Jordan and Brayton 1878)-none.
Nominal forms of Notropis definitely or likely based on hybrids of the genus
1. Opsopoeodus borealis Harper and Nichols 1919-possibly based on a hybrid, one parent
of which is presumed to be Notropis hudsonius.
2. Notropis germanus Hay 1887-Hybognathus hankinsoni x Notropis heterolepis (see Hubbs
1951b).
1978
BULLETIN FLORIDA STATE MUSEUM
3. Notropis kanawha Jordan and Jenkins 1889-Notropis rubellus x Notropis rolucellus (see
Bailey and Gilbert 1960)
4. Notropis macdonaldi Jordan and Jenkins 1889-Notropis cornutus X Notropis rubellus
(see Hubbs and Moore 1940)
5. Notropis umbrifer Hay 1887-likely based on a hybrid, one parent of which may be No-
tropis topeka.
NOMINAL FORM OF Notropis BASED ON HYBRIDS
OF OTHER GENERA
1. Notropis henryi Fowler 1942-Clinostomus funduloides X Nocomis leptorephalus (see
Gilbert 1978).
Nomina nuda OF Notropis
1. Notropis comalis Jordan and Gilbert 1885-name listed by Jordan (1885b:812); most likely
based on Notropis stramineus, but possibly could refer to Notropis buchanani or
Notropis amnis.
2. Notropis ionthas Jordan and Gilbert 1886-described as a supposed junior synonym of No-
tropis dilectus (=N. atherinoides) (Jordan and Gilbert 1886:11), but name is un-
available since it was proposed as a junior synonym and was not treated as an avail-
able name, with its original date and authorship, prior to 1961 (International Code
of Zoological Nomenclature [1964: Articles 11(d) and 16(b) (ii)]); based on Notropis
perpallidus Hubbs and Black (see Snelson and Jenkins 1973:293).
3. Notropis ochoterenai Hubbs and Gordon, in Hubbs 1937-based on Notropis aquirrepequenoi
Contreras and Rivera 1973.
4. Notropis xaenocephalus octoradius Baughman 1950-name indicated as having first ap-
peared in a mimeographed key to Texas fishes by G. H. Soulen (1941); based on
Notropis chalybaeus (Cope) (C. C. Swift, in litt.).
5. Notropis spilurus Gilbert and Swain 1885-name listed by Jordan (1885:814); based on
specimens of Notropis stilbius (Jordan) with a conspicuous caudal spot (see Gilbert
1891:154).
NOMINAL FORMS OF Notropis Now REFERRED TO OTHER GENERA
1. Notropis celayensis Alvarez 1958-a junior synonym of Yuriria alta (Jordan 1880) (R. R.
Miller, in litt.).
2. Notropis ipni Alvarez and Navarro 1953-a valid species now referred to the genus Dionda
(see Hubbs and Miller 1974:1).
3. Notropis rasconis Jordan and Snyder 1899-a valid species now referred to the genus Dionda
(see Hubbs and Miller 1974:1).
ACCOUNTS OF FORMS REFERRED TO Notropis
Notropis chalybaeus abbotti Fowler 1904
Fowler 1904a:239-240, pl. 17 (upper fig.). TYPE LOCALITY: broken dam on Batsto R., Burling-
ton County, New Jersey. HOLOTYPE: ANSP 19860 (45.0), E. D. Cope. PARATYPES: ANSP 19861-
19878 (18), paratopotypes.
REMARIS.-Fowler (1910: pl. 19, fig. 40) illustrated the holotype. Anal-ray count for holotype
8. Original description date 7 April 1904.
= Notropis chalybaeus (Cope)
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
Alburnus acutus Lapham 1854
Lapham 1854:101, figs. 1-2. TYPE LOCALITY: lower part of Milwaukee R. (below first dam),
Milwaukee Co., Wisconsin. TYPES: None located.
REMARKS.-Original description reproduced in Hubbs (1945:16). Hubbs and Lagler (1958:81)
regarded acutus as a valid subspecies of N. atherinoides, but the systematics of this species have
not yet been adequately worked out. Original description date 1 April 1854.
= Notropis atherinoides Rafinesque
Notropis aeneolus Hay 1887
Hay 1887:245-246. TYPE LOCALITY: Saline R., ca. 5 mi. N of WaKeeney, Trego Co., Kansas.
HOLOTYPE: USNM 37945 (56.5), M. J. Thompson, July 1885.
REMARKs.-Jordan and Evermann (1896:266) were first to synonymize this form with N. to-
peka. Original description date 2 July 1887.
= Notropis topeka (Gilbert)
Notropis aguirrepequenoi Contreras and Rivera 1973
Contreras and Rivera 1973:9-23, fig. 1. TYPE LOCALITY: Rio Pilon (trib. to Rio Soto la Ma-
rina), at crossing of hwy. 85, town of Mainero, Tamaulipas, Mexico. HOLOTYPE: UNAM IB/CML-
P434 (39.7), S. Contreras, D. Molina, A. Aseff, and A. Martinez, 21 May 1966. PARATYPES: UANL
1137 (416), IPN P-3678 (4), UNAM IB/CML-P435 (4), TU 94116 (20), UMMZ 197492 (4), USNM
214998 (4) (all ex UANL 1137), paratopotypes; UANL 290 (1), UANL 297 (109), UANL 1141 (150).
REMARKS.-This species was earlier listed by Hubbs (1937) as a nomen nudum, Notropis ocho-
terenai. Original description date 5 October 1973.
= Notropis aguirrepequenoi Contreras and Rivera
Notropis alabamae Jordan and Meek 1884
Jordan and Meek 1884:476-477. TYPE LOCALITY: trib. to Alabama R., Montgomery, Alabama.
LECTOTYPE: USNM 35297 (44.5), Col. Marshall McDonald. Snelson (1972:21) designated lecto-
type. LECTOPARATYPES: USNM 203323 (5), paratopotypes.
REMARKS.-Jordan and Evermann (1896:298) erroneously listed USNM 35295 as a type num-
ber. Species first synonymized with N. lirus (Jordan, 1885b:815), but subsequently synonymized
with N. bellus by Smith-Vaniz (1968:43). Original description date 23 October 1884.
= Notropis bellus bellus (Hay)
Notropis megalops albeolus Jordan 1889
Jordan 1889b:123. TYPE LOCALITY: Roanoke R., near Roanoke, Virginia. LECTOTYPE: USNM
40177 (76.0), D. S. Jordan and B. W. Evermann, 31 July-2 August 1888. Gilbert (1964:152) desig-
nated lectotype. LECTOPARATYPES: USNM 177840 (14), CAS-SU 611 (6), paratopotypes; MCZ
31984 (1), paratopotype (?).
REMARKS.-Gilbert (1964:151-157) reviewed species. Original description date (31 December)
1889.
= Notropis albeolus Jordan
Notropis albeolus Eigenmann and Eigenmann 1893
Eigenmann and Eigenmann 1893a:152. TYPE LOCALITY: S. Saskatchewan R., Medicine Hat,
Alberta, Canada. HOLOTYPE: BMNH 1892.12.30.531 (53.5), C. H. Eigenmann, August or Sep-
tember 1892.
REMARKS.-As the species name albeolus was preoccupied in Notropis, the name jordanii was
later substituted by Eigenmann and Eigenmann (1893b). Hubbs (1926:41) first synonymized this
form with N. heterolepis, but later reidentified it as N. blennius jejunus following examination
of holotype (Hubbs, in litt.). The type has 1,4-4,2 pharyngeal teeth (left arch deformed) and seven
anal rays. Original description gives type locality as "Medicine Hat, Assiniboia," but the former
locality is in Alberta and the latter in Saskatchewan. Only one specimen formed basis for original
1978
BULLETIN FLORIDA STATE MUSEUM
description, and the label in the holotype bottle shows Medicine Hat to be the correct type lo-
cality. Original description date 4 February 1893.
= Notropis blennius (Girard)
Notropis alborus Hubbs and Raney 1947
Hubbs and Raney 1947:1-17, pl. 1 (figs. 1-2). TYPE LOCALITY: Brush Cr., trib. to Deep R., 5
mi. W of Siler City, Randolph Co., North Carolina. HOLOTYPE: UMMZ 138489 (42.0), E. C.
Raney and E. A. Lachner, 6 March 1940. PARATYPES: UMMZ 138490 (19), paratopotypes; UMMZ
132797 (1), UMMZ 138332 (8), UMMZ 138333 (3), UMMZ 138488 (8), USNM 40347 (27), USNM
93227 (16), USNM 107615 (48), CU 3538 (5), CU 9619 (2).
REMARKS.-Original description date 25 February 1947.
= Notropis alborus Hubbs and Raney
Notropis bellus alegnotus Snelson 1972
Snelson 1972:37-43; figs. 3a, 9. TYPE LOCALITY: Five Mile Cr., trib. to Valley Cr., U.S. hwy.
11 bridge, 5.0 air mi. SW of jet. of st. hwy. 150 and U.S. hwy. 11 in Bessemer, T19S, R5W, Sec.
36, Jefferson Co., Alabama. HOLOTYPE: CU 53346 (42.3), F. F. Snelson, Jr. and B. W. Menzel,
19 May 1969 (orig. no. FFS 69-7). PARATYPES: CU 53354 (68), TU 58810 (10), UMMZ 197674
(10), USNM 204313 (10), paratopotypes.
REMARKS.-Original description date 30 November 1972.
= Notropis bellus alegnotus Snelson
Hudsonius aletes Jordan and Evermann 1927
Jordan and Evermann 1927:502. TYPE LOCALITY: Switz City Swamp, Green County, Indiana.
TYPES: None. Based on three specimens, collected by C. H. Gilbert in late August, 1883 (and
identified by him as Notropis heterodon [Gilbert 1884b:207-208]), which were catalogued into
the Stanford University collection as SU 2580. Inasmuch as these were destroyed as a result of
the 1906 earthquake (C. C. Swift, in lift.), the original description evidently was based on ma-
terial no longer extant.
REMARKS.-Gerking (1945:60) regarded this form as a subspecies of N. xaenocephalus (auct.).
Bailey, Winn, and Smith (1954:126) were first to synonymize it with N. roses (=N. texanus).
Original description date 27 April 1927.
= Notropis texanus (Girard)
Alburnellus altipinnis Cope 1870
Cope 1870:464-465. TYPE LOCALITY: Yadkin R., Roane (= Rowan) Co., North Carolina. LEC-
TOTYPE: ANSP 2846 (41.3), E. D. Cope, Fall 1869. Fowler (1910: pl. 19, fig. 37) designated lec-
totype by illustrating "type" (ANSP 2846); calculated length 41.3 mm SL. LECTOPARATYPE:
ANSP 2847 (1), paratopotype. The third syntype (ANSP 2031) has been reidentified as N. peter-
soni, and is recatalogued under that name.
REMARKS.-Condition of ANSP 2846-2847 fair; condition of ANSP 2031 poor. Counts for
ANSP 2846: teeth 2,4-4,1 (lesser tooth in right arch represented only by a hole); anal rays 9.
Counts for ANSP 2847: teeth 1,4-4,1; anal rays 9. Counts for ANSP 2031 (N. petersoni): teeth
2,4-4,2; anal rays 7. Hubbs and Raney (1948) reviewed species and also described several new
subspecies. Original description date 21 November 1870.
= Notropis altipinnis (Cope)
Alburnus amabilis Girard 1857
Girard 1857:193. TYPE LOCALITY: Rio Leona(trib. to Rio Nueces), Uvalde, Uvalde Co., Texas.
SYNTYPES: USNM 72 (26, 42.8-50.2; one right pharyngeal arch in bone collection), ANSP 3151
(ex USNM 72) (1,45.5), MCZ 1684 (ex USNM 72) (2, 45.5-48.5), J. H. Clark, 1851.
REMARKS.-Girard (1859a:52; pl. 29, figs. 10-13) listed only 15 original syntypes and illus-
trated one of these. Types in relatively good condition for old specimens. Counts for USNM syn-
types: teeth 2,4-4,1 (1); anal rays 8 (2), 9 (22), 10 (2); lateral-line scales 36 (1). Counts for MCZ
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
syntypes: teeth4,2 and 2,4-4,2: anal rays 9 in both: predorsal scales 17 and 18: bodv-circnmfer-
ential scales 11-2-9= 22 in both; caudal-peduncle circumferential scales 5-2-5= 12 in both: lateral-
line scales 36 and 37. Counts for ANSP svntvpes: teeth 2,4-4,1: anal rays 9.
As can be seen from the master species list (p. 17), several nominal forms described by
Girard (1857) are identical with N. amabilis, the original description of which also appeared in
the same paper. Jordan and Gilbert (1883:195) svnonvmized Alburnus socios with A. imegalops,
although Jordan (1885a:123) subsequently refuted this and regarded the two as distinct. Jordan
and Gilbert (1883:177) also svnonvmized Cyprinella luxiloides with C. mnarrostomia. The species
amabilis was regarded as valid in both the above papers, as it was in all subsequent papers in
which the name appeared (Evermann and Kendall 1894:102; Jordan and Evermann 1896:292;
Fowler 1910:288; Jordan, Evermann, and Clark 1930:124). Baughman (1950:130) listed N. anta-
bilis and N. swaini as valid, but made no comment regarding the other forms; and Hubbs, Kuehne,
and Ball (1953:226) indicated that specimens recorded as V. swaini by Jordan and Gilbert
(1886:24) and Evermann (1892:79) from the Guadelupe River system are, m fact, V. amabilis.
Miller (1976:11) synonymized C. macrostoima with N. iamabilis and, as first reviser, selected the
latter name. Other than this, none of the above forms have been identified or placed in synonvmy.
I conclude, based on examination of the descriptions, figures, and extant type material, that A/-
burnus miegalops (including the replacement name Xotropis srwaini), Alburnuss socius, and (prob-
ably) Cyprinella luxiloides are also identical to .V. amabilis. and, as first reviser, I formally select
the last name as the one to be used. Original description date 25 April 1857.
= Notropis amlabilis (Girard)
Hudsonius aImanrus Girard 1857
Girard 1857:210. TYPE LOCALITY: Potomac R.. between Chesapeake Bay and Washington,
D.C. SYNTYPES: USNM 15 (one set of pharyngeal arches), USNM 16 (one set of arches), USNM
17 (four sets of arches).
REMARKS.-Counts for pharyngeal arches: USNM 15 (?-4,2), USNM 16 (?-4,1), USNM 17
(2,4-4,2 in two; 1,4-4,1 in one; 1,4-4,0 in one). Jordan (1876b:281) was first to synonvmize this
form with N. hudsonius. Original description date 25 April 1857.
= Votropis hudsonius (Clinton)
Rutilus amblops Rafinesque 1820.
See pages 21, 90.
Notropis amniis amniis Hubbs and Greene 1951
Hubbs and Greene, in Hubbs 1951a:2-14, pl. 1 (figs. 1-2). TYPE LoC:ALITY: Mississippi R., 1
mi. N of Prairie du Chien, Crawford Co., Wisconsin. HOLOTYPE: UMMZ 75435 (43.0), L. P.
Schultz and C. M. Tarzwell, 27 August 1928. PARATYPES: UMMZ 78246 (68), MCZ 36079 (ex
UMMZ 78246) (3), paratopotypes; UMMZ 72(04 (2), UMMZ 72022 (1), UMMZ 76634 (3), UMMZ
76797 (2), UMMZ 77705 (8), UMMZ 77959 (1), UMMZ 78186 (30) UMMZ 78222 (16), USNM
117559 (ex UMMZ 78222) (2), UMMZ 78279 (11), UMMZ 78312 (11), UMMZ 78361 (8), UMMZ
100918 (2), UMMZ 100949 (4).
REMARKS.--Species name ainnis first appeared as a nonimn nuduni in Greene (1935:96-97). For
further remarks on the nomenclature of this species see accounts of .. nmocomis Jordan and Gil-
hert and N. nocomis Evermann. Clemmer (1970) reviewed species. Original description date 14
March 1951.
= Votropis ainis Hulbbs and Greene
Alburnerllus uamonu s Abbott 1874
Abbott 1874:334-335, fig. 78. TYE l.o( LITY: Delaware and Raritan Canal, probably in
Mercer Co., New Jersey. TYPES: None located. Two specimens, 66.4-73.6 mm SI (MCZ 30697),
are catalogued as "types," but Snelson (1968:779-780), on basis of locality and other accompany-
ing data, feels they should not be so considered.
1978
BULLETIN FLORIDA STATE MUSEUM
REMARKs.-Snelson (1968) reviewed species. Original description date (30) June 1874.
= Notropis amoenus (Abbott)
Cyprinella analostana Girard 1859
Girard 1859b:58-59. TYPE LOCALITY: Rock Cr., trib. to Potomac R., Washington, D.C. TYPE:
None located.
REMARKS.-Gibbs (1963) reviewed species. Form very close to N. whipplei. Original descrip-
tion date (31 December) 1859.
= Notropis analostanus (Girard)
Notropis anogenius Forbes 1885
Forbes 1885:138-139. TYPE LOCALITY: Fox R., McHenry, McHenrv Co., Illinois. LECTOTYPE:
INHS 26948 (43.0), S. H. Forbes (presumably), 8-10 May 1880. Bailey (1959:120) designated
lectotype. LECTOPARATYPES: INHS 26949 (5), USNM 64153 (7), paratopotypes.
REMARKs.-Lectotvpe in poor condition (desiccated). Bailey (1959) reviewed species. Original
description date (31) March 1885.
= Notropis anogenus Forbes
Notropis telhscopus arcransanus Meek 1891
Meek 1891:1.33-134. TYPE LOCALITY: Spring Branch, town of Mammoth Spring, Fulton Co.,
Arkansas. LECTOTYPE: CAS-SU 1027 (50.6), S. E. Meek, L. Rettger, and F. J. Drew, 5 August
1889. Lectotype herein designated. LECTOPAHATYPES: CAS-SU 68040 (4), paratopotypes: ANSP
3251-3258 (8).
REMARKs.-Although Meek (1891:134) took his original specimens at Mammoth Spring, he
also mentions having examined material collected by D. S. Jordan et al from the White River,
near Eureka Springs, Carroll County, Arkansas, in September 1884. Of the two extant type series,
one (CAS-SU 1027) is from Mammoth Spring, and the other (ANSP 3251-3258) is from near
Eureka Springs. Inasmuch as syntypes of N. telescopes arransanus exist from both places, and
the type locality has, in consequence, never been clearly specified; a lectotype is herein selected
in order to stabilize this situation. Gilbert (1969:489-490) synonymized this form with N. tele-
scopus. Lectotype and CAS-SU lectoparatypes in poor condition. Anal-ray counts for ANSP
lectoparatypes 10 (3), 11 (5). Original description date 19 June 1891.
= Notropis telescopes (Cope)
Leuiriscus gardens Cope 1868
Cope, in Ginther 1868:257-258. TYPE LO( ALITY: headwaters of Roanoke R., Montgomery
Co., Virginia. LECTOTYPE: ANSP 3268 (66.0), E. D. Cope, July 1867. Fowler (1910: pl. 21, fig.
56) designated lectotype by illustrating "type" (ANSP 3268); calculated length 67.9 mm SL.
LECTOPARATYPES: ANSP 3269-3315 (48), BMNH 1868.1.10.23 (2), paratopotypes. According to
Giinther (1868), eight BMNH syntypes were originally present.
REMARKs.-F. F. Snelson (Ms.) is reviewing species. Original description date 14 March 1868.
= Notropis gardens (Cope)
Alhmirncllus large Cope 1869
Cope 1869:388. TYPE LOCALITY: either Detroit or St. Josephs River, Michigan. TYPES: None
located.
REMARKS.- lHbbs (1926:45) was first to synonymize this form with Notropis photogenis. Orig-
inal description date (31) August 1869.
= Notropis photogenis (Cope)
Plargyrus argentatus Girard 1857
Girard 1857:212. TYPE LO:ALITY: James River system, Virginia. TYPES: None located.
REMARKS.-Jordan and Gilbert (1883:187) were first to synonymize this form with N. cornutus.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
Original description date 25 April 1857.
= Notropis cornutus (Mitchill)
Photogenis ariommnus Cope 1868
Cope 1868a:164-165. TYPE LOCALITY: White R., near Indianapolis, Marion Co., Indiana.
HOLOTYPE: ANSP 16488 (54.0), W. P. Clark, 1865 or 1866.
REMARKs.-Gilbert (1969) reviewed species. Fowler (1910: pl. 20, fig. 43) illustrated "type,"
calculated length 54.9 mm SL. Most literature records of "Notropis arionimus" between 1939-
1969 actually refer to N. telescopus. Original description date 11 May 1868.
= Notropis ariormmus (Cope)
Notropis asperifrons Suttkus and Ranev 1955
Suttkus and Ranev 1955c:3-33, figs. 1-2. TYPE LOCALITY: Holly Cr.. Ramhurst, 8 mi. N of
Murray Co. line, on U.S. hwy. 411, Murray Co., Georgia. HOLOTYPE: CU 28262 (50.0), R. H.
Gibbs and P. P. Caswell, 12 June 1952. PARATYPES: CU 2826:3 (7), paratopotypes; CU 28261 (1),
CU 28260 (3), TU 4251 (28), TU 3426 (8), TU 2974 (22), TU 3063 (5), UMMZ 111122 (2), UMMZ
111125 (7), UMMZ 162594 (2), USNM 164968 (1), USNM 164969 (1), UMMZ 139104 (3).
REMARKS.-Swift (1970) reviewed species. Original description date 8 July 1955.
= Notropis asperifrons Suttkus and Ranev
Notropis atherinoides Rafinesque 1818
Rafinesque 1818:204. TYPE LOCALITY: "Lake Erie." TYPES: None located. Specimens said
to have been collected bv DeWitt Clinton and deposited in Lyveum of Natural History.
REMARKs.-Type species of genus Notropis. Original description date (31) January 1818.
= Notropis atherinoides Rafinesque
Notropis atrapiculus Snelson 1972
Snelson 1972:58-71, figs. 3c, 4e-h. TYPE LOCALITY: Sikes Cr., trih. to West Fk. of Chocta-
whatchee R., on unnumbered county rd., 3.5 air mi. ENE of center of Clio, T9N, R25E, Sec. 33,
Barbour Co., Alabama. HOLOTYPE: CU 53343 (47.7) (adult tuberculate male), F. F. Snelson, Jr.
and B. W. Menzel, 25 May 1969 (orig. no. FFS 69-18). PARATYPES: CU 53351 (20), CU 53142
(26), USNM 204314 (8), paratopotvpes: TU 2509 (36).
REMARKS.-Earlier literature records of N. roseipinnis are based in part on this species.
Original description date 30 November 1972.
= Notropis atrapiculu s Snelson
Lythrurns stripes Jordan 1878
Jordan 1878b:59. TYPE LOCALITY: "Various streams in Union and Johnson countries, Illinois."
Locality on label of lectotype indicated as Cache R., Johnson Co., Illinois. LECTOTYPE: USNM
26295 (46.0), S. A. Forbes, Snelson and Pflieger (1975:235) designated lectotype. LECTOPARA-
TYPES: CAS-SU 1989 (2), "Illinois."
REMARKS.-Form regarded as subspecies of N. gardens by Jordan (18851:814) and as subspecies
of N. umbratilis by Jordan and Evermann (1896:300). Original description date (30) June 1878.
= Notropis umbratilis cynocephalus (Copeland)
Notropis crayuga atrocaldalis Evermann 1892
Evermann 1892:76. TYPE LOCALITY: Neches R., ca. 14 mi. E of Palestine, Anderson Co., Texas
(at Palestine and Rusk RR bridge). LECTOTYPE: USNM 45557 (39.6), B. W. Evermann, J. T.
Scovell, R. R. Gurley, and J. A. Singley, 24 November 1891. Evermann and Kendall (1894:
pl. 16) designated lectotype by illustrating "type." Label in type jar says "specimen drawn."
LECTOPARATYPES: USNM 125171 (14), CAS-SU 2139 (1), paratopotypes.
1978
BULLETIN FLORIDA STATE MUSEUM
REMARKs.-Hubbs and Ortenhurger (1929b:67) considered this possibly to be a distinct spe-
cies, but Hubbs (1951a:13) was first actually to validate it. Original description date 25 May
1892.
= Notropis atrocaudalis Evermann
Moniana aturata Girard 1857
Girard 1857:200. TYPE LOCALITY: Pinto Creek, "Piedra Pinta," 15-20 mi. SE of Del Rio,
Kinney Co., Texas. (Not "Piedra Painte," New Mexico.) SYNTYPES: USNM 118 (43, 33.5-55.0,
plus one left arch in bone collection) (in two jars [2+41 specimens]), USNM 125084 (4, 42.7-
49.3), MCZ 1689 (ex USNM 118) (2, 48.0-48.3), ANSP 2833-2834 (ex USNM 118) (2, 40.2-45.5),
J. H. Clark, 1851.
REMARKS.-Girard (1859a:56; pl. 30, figs. 13-16) listed only 20 original syntypes and illustrated
one of these. Jordan (1885a:125) was first to synonymize this form with N. proserpinus, an action
confirmed by Hubbs and Ortenburger (1929b:75) following examination of types. The two MCZ
syntypes, in relatively good condition, are high tuberculate males tubercless in front of eye large),
both with 4-4 pharyngeal teeth and 8 anal rays. The two ANSP svntypes, also in relatively good
condition, have 4-4 teeth and 8 anal rays. The type locality is presumed to be that listed as "Piedra
Pinta" in the table of distances of Emory's (1857:135) report. No locality with this name appears
for the "New Mexico" section of the table, and the remaining data accompanying the specimens
(collector and year) are in accord with this interpretation. Original description date 25 April
1857.
= Notropis prose'rpinus (Girard)
Notropis aztecus Woolman 1894
Woolman 1894:63-64; pl. 2, upper right fig. TYPE LOCALITY: canal in Mexico City, D.F.,
Mexico. SYNTYPES: USNM 45569 (1, 63.0), USNM 47505 (1, 58.7), USNM 125172 (1, 57.5), CAS-
SU 644 (10, 45.6-73.7), FMNH 6604 (11, 42.0-77.3), BMNH 1894.1.27.58-61 (4, 56.0-69.5), A. J.
Woolman and U. O. Cox, August 1891.
REMARKS.-Jordan and Evermann (1896:258) synonymized this form with Codoma tittata
Girard 1857, but pointed out that the latter was preoccupied in Notropis by Leuc'iscus cittattus
DeKay 1842. Jordan and Evermann (19(): pl. 45, fig. 122) and later Meek (1904:60, fig. 1:3) illus-
trated one of the syntypes (USNM 45569). Miller (1976:10) tentatively called Ceratichthys sallaei
Gfinther 1868 a senior synonym of N. aztecus. Counts for USNM 45569: anal rays 7; lateral-line
scales 45. Counts for USNM 47505: anal rays 7; lateral-line scales 46. Counts for USNM 125172:
anal rays 7; lateral-line scales 50; pharyngeal teeth 4-. Original description date 3 May 1894.
= Notropis sallei (Giinther)
Notropis haileyi Suttkns and Raney 1955
Suttkus and Raney 1955a:71-86, figs. 1-2. TYPE LOCALITY: Sawacklahatchee Cr., 1.7 mi. \V of
Society Hill, on U.S. hwy. 80, Macon Co., Alabama. HOLOTYPE: CU 28224 (59.6), R. D. Suttkus,
C. F. Cole, and R. H. Gibbs, Jr., 12 June 1949. PARATYPES: CU 16020 (66), paratopotypes; TU
3195 (4), TU 2637 (30), TU 3430 (75), TU 2965 (4), TU 3066 (41), USNM 163568 (1), UMMZ
105526 (2), UMMZ 105554 (73), UMMZ 124018 (1), UMMZ 124024 (1), UMMZ 124094 (41),
UMMZ 124117 (45), UMMZ 146517 (5), UMMZ 146533 (16), UMMZ 146544 (97).
REMARKS.-Swift (1970) reviewed species. Original description date 12 January 1955.
= Notropis haileyi Suttkus and Raney
Notropis bairdi Hubbs and Ortenburger 1929
Hubbs and Ortenburger 1929a:29-32. TYPE LOCALITY: Red R., 6-9 mi. SW of Hollis, Harmon
Co., Oklahoma. HOLOTYPE: UMMZ 80346 (64.0), A. I. Ortenburger et al., 16 June 1926. PARA-
TYPES: UMMZ 80344 (1), UMMZ 80345 (590 originally, now 574), USNM 93428 (1), USNM
117560 (10), BMNH 1933.1.24.3-5 (3), and OAM uncat. (2) (all four series ex UMMZ 80345),
UOMZ 6350 (13(X)), paratopotypes; UMMZ 73031 (9), UMMZ 80368 (118), UMMZ 80378
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
(934 originally, now 929), MCZ 36084 (ex UMMZ 80378) (5), UMMZ 80387 (102 originally, now
94), UF 14599 (ex UMMZ 80387) (8), UMMZ 80402 (3), UMMZ 80409 (1), UOMZ 6183 (140),
UOMZ 6197 (3), UOMZ 6212 (1), UOMZ 6284 (100 ), UOMZ 6295 (100).
REMARKS.-Original description date 16 March 1929.
= Notropis bairdi Hubbs and Ortenburger
Cyprinella beckwithi Girard 1857
Girard 1857:197. TYPE LOCALITY: sluices of Arkansas R., near Ft. Makee, Kansas. (Said to be
"Ft. Walker" in ANSP catalogue). SYNTYPES: MCZ 1801 (1, 40.3), USNM 135 (two left and one
right pharyngeal arches in bone collection), Lt. E. G. Beckwith, July 1853. The one syntype
(ANSP 3053) Fowler (1910:280) said was "nearly dissolved" was not located.
REMARKS.-Girard (1858:268) listed nine original syntypes. Jordan (1885a:123) synonymized
this form with N. bubalinus, and Hubbs and Ortenburger (1929b:75) later synonymized both
with N. lutrensis. MCZ syntype contorted but otherwise in relatively good condition; specimen
examined by Hubbs (see Hubbs and Ortenburger 1929b:75). Counts for MCZ syntype: teeth
1,4-4,1 (Hubbs said 2,4-4,0); anal ravs 9. Counts for USNM syntypes pharyngeall arches): left
arches 1,4- and 0,4-; right arch -4,1 (hole where lesser tooth should be). Original description date
25 April 1857.
= Notropis lutrensis (Baird and Girard)
Minnilus bells Hav 1881
Hay 1881:510-511. TYPE LOCALI TY: Catawba Cr. and tributaries, Artesia, Lowndes Co., Mis-
sissippi. LECTOTYPE: USNM 27426 (44.0), O. P. Hay, March-April 1880. Snelson (1972:20) desig-
nated lectotype. LECTOPARATYPES: USNM 203332 (ex USNM 27426) (3). CAS-SU 756 (6), CAS-
SU 2526 (1), paratopotvpes.
REMARKs.-Originally 40 svntvpes were extant, of which 36 were from the above locality and
4 were from the Noxubee R., Macon, Noxubee Co., Mississippi. Only the types listed above are
now accounted for. Snelson (1972:20-37) reviewed species. Original description date 23 Febru-
ary 1881.
= Notropis hellus bells (Hay)
Hybopsis bifrenatus Cope 1869
Cope 1869:384-385. TYPE LO(ALITY: Trib. of Schuylkill R., Conshohocken, Montgomery Co.,
Pennsylvania. TYPES: None located. In the original description Cope mentioned having collected
nearly 100 specimens. Fowler (1909:531) stated, "Cope's types of this species (Hybopsis bi-
frenatus) seem to be lost."
REMARKs.-Harrington (1947) mapped distribution and Jenkins and Zorach (1970) reviewed
species. Original description date (31) August 1869.
= Notropis bifrenatus (Cope)
Cyprinella billingsiana Cope 1871
Cope 1871:439. TYPE LOCALITY: Missouri R., St. Joseph, Missouri. LECTOTYPE: ANSP 2952
(34.5), Dr. William Stimpson. Fowler (1910: pl. 17, fig. 19) designated lectotype by illustrating
"type" (ANSP 2952); calculated length 35.0 mm SL. LECTOPARATYPES: ANSP 2953-2991 (39),
paratopotypes.
REMARKS.-Jordan (1885b:812) was first to synonymize this form with N. lutrensis. Condition
of syntypes soft, but otherwise fairly good. Largest specimen with teeth 4-?; anal rays 9. Counts
for other specimens: teeth 4-4 (2), 1,4-4,1 (2); anal rays 8 (3), 9 (36). Original description date
(31 December) 1871.
= Notropis lutrensis (Baird and Girard)
Notropis hltrensis blairi Hubbs 1940
Hubbs 1940:6-8. TYPE LOCALITY: Garden Springs (= Monument Spring), trib. to Pefia Colo-
BULLETIN FLORIDA STATE MUSEUM
rado Cr. (Maravillas Creek drainage), 12-13 mi. SSW of Marathon, Brewster Co., Texas. Eleva-
tion 3700 ft. HOLOTYPE: UMMZ 104170 (44.0), Josselyn Van Tyne and W. F. Blair, 16 April 1937.
PARATYPES: UMMZ 104171 (15), UMMZ 127335 (22), BU (uncat.) (ex UMMZ 127335) (3), para-
topotypes (UMMZ 127335 collected subsequent to UMMZ 104171); UMMZ 120330 (10).
REMARKs.-Contreras (1975) tentatively synonymized this form with the typical subspecies.
Original description date (31) October 1940.
= Notropis lutrensis (Baird and Girard)
Alburnops blennius Girard 1857
Girard 1857:194. TYPE LOCALITY: Arkansas R., near Fort Smith, Arkansas. LECTOTYPE: USNM
67 (55.9), Dr. George F. Shumard, 1853. Suttkus (1958:308) designated lectotype. LECTOPARA-
TYPES: USNM 171791 (ex USNM 67) (5), USNM 64 (one pair of pharyngeal arches in bone col-
lection), MCZ 1784 (1), ANSP 3030 (1), paratopotypes.
REMARKS.-Girard (1858:261; pl. 57, figs. 13-16) listed 18 original syntypes and illustrated
one of these. Hubbs, in 1958, identified syntype in British Museum (BMNH 1883.12.14.213) as
N. colucellus. Pharyngeal-tooth count for USNM 64:2,4-4,2. Counts for MCZ 1784: teeth 1,4-4,1;
anal rays 7. Counts for ANSP 3030: teeth 2,4-4,2; anal rays 7. The specific name blennius was long
incorrectly applied to the species now called N. stramineus (Jordan and Evermann 1896:261-
262), until Hubbs (1926:42-43) clarified its status. Hubbs and Bonham (1951:104) recognized two
subspecies of N. blennius (the nominate southern subspecies and a northern subspecies, jejunus),
but these are not substantiated by the preliminary findings of Suttkus and Clemmer (1968). Orig-
inal description date 25 April 1857.
= Notropis blennius (Girard)
Opsopoeodus bollmani Gilbert 1890
Gilbert 1890:226-227. TYPE LOCALITY: lagoon of Buckhead Cr., 1 mi. SW of Millen, Jenkins
Co., Georgia. SYNTYPES: USNM 61570 (4, 33.8-47.4), C. H. Bollman and B. Fesler, late June 1889.
REMARKS.-Bailey, Winn, and Smith (1954:129) were first to synonymize this form with N.
maculatus. Original description date (31 December) 1890.
= Notropis maculatus (Hay)
Notropis hoops Gilbert 1884
Gilbert 1884a:201-202. TYPE LOCALITY: Salt Cr., Brown Co., Indiana; also Flat Rock Cr.,
Rush Co., Indiana. SYNTYPES: USNM 34982 (3, 44.0-45.6), MCZ 35961 (1, 54.3), C. H. Gilbert
et al. CAS-SU 3794 (20, 47.4-61.0 [plus one specimen of Hybopsis amblops]); probable syntypes
(see below).
REMARKS.-Originally 10 syntypes were listed for USNM 34982, all from Salt Creek. The four
specimens still remaining, including the one at the MCZ, leave six syntypes from this series un-
accounted for. In the original description Gilbert (1884a:202) also listed 30 syntypes from Flat
Rock Creek, collected by W. P. Shannon. The California Academy of Sciences collection (CAS-
SU 3794) has 20 specimens of N. hoops (plus one specimen of Hybopsis amblops) from the Flat
Rock Creek locality. These probably are syntypes, though C. H. Gilbert is listed as the collector,
and nothing in the jar indicates that these are type specimens. Because of the doubt regarding the
status of these specimens, I recommend that a lectotype, if selected, be chosen from among the
USNM or MCZ types. Counts for USNM and MCZ types: teeth 1,4-4,1, in all; anal rays 8 in all.
Original description date 1 September 1884.
= Notropis boops Gilbert
Opsopoeodus borealis Harper and Nichols 1919
Harper and Nichols 1919:266. TYPE LOCALITY: Lake Athabaska, Ft. Chipewvan, Alberta,
Canada. HOLOTYPE: NMC 1048 (27.0), F. Harper, 15 June 1914. Type lost, according to McAllis-
ter (1965:10); this reconfirmed by McAllister (in litt., 6 August 1971).
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
REMARKS.-Exact status uncertain. Possibly a junior synonym of X. hudlsoniuls, as suggested
by Kendall (1924) and by Gilbert and Bailey (1972:14), this based on geographic locality and the
"sharply defined black blotch, about half diameter of eye, at base of caudal, partly on the fin."
However, the pharyngeal tooth count was given as 4-, and dorsal- and anal-fin ray counts as 9
each, all of which differ from what one would expect in N. hudsonius from western Canada. Pos-
sibly this nominal form is based on a hybrid, one parent of which is N. hudsonius. Original descrip-
tion date 22 September 1919.
= Notropis hudsonius (Clinton) or possibly a hybrid involving that species.
Leuciscus ourcardi Giinther 1868
Giinther 1868:485. TYPE LOCALITY: Cuernavaca, Mexico. SYNTYPES: BMNIt 1868.3.3.4-7
(4, 61.0-72.0), M. Boucard.
REMARKS.-This species has been assigned to the genus Hyhopsis, based on the presence of a
barbel at each corner of the mouth, but Cortes (1968) recently placed it in Notropis. Leuciscus
boucardi, Graodus nigrotaeniatus, and Ceratichthys cumingii, all of which were originally de-
scribed by Giinther (1868), are synonyms. Meek (1904:68) was first to point out the relationships
of the first two forms, and, as first reviser, selected the name houcardi. Carl L. Hubbs examined
the types of C. cumingii in the British Museum in 1958, and synonvmization of that form with
N. boucardi is based on his analysis. Original description date 14 March 1868.
= Notropis boucardi (Giinther)
Plargyrus bourimoni Girard 1857
Girard 1857:196. TYPE LOCALITY: Sweetwater R., Nebraska. HOLOTYPE: ANSP 3236 (59.5). J.
Soul6 Bowman. This may not be the type (see below).
REMARKS.-Girard (1858: pl. 59, figs. 1-5) illustrated the unique type (USNM 65), which
measures 51 mm SL and was said to be drawn to actual size. The length of the ANSP individual
(60.5 mm SL) raises doubt that this is the type specimen. Also, as the type is unique, it seems un-
likely that the USNM would have sent it to the ANSP. Jordan and Gilbert (1883:187) were first
to synonymize this form with N. cornutus. Condition of type (?) fairly good, considering age.
Original description date 25 April 1857.
= Notropis cornitus (Mitchill)
Notropis braytoni Jordan and Evermann 1896
Jordan and Evermann 1896:264-265. TYPE LOCALITY: Rio Monterrev (trib. to Rio San Juan),
Cadereita (= Cadereyta), Nuevo Leon, Mexico. SYNTYPES: USNM 39657 (2, 47.5-52.4), Lt. D. N.
Couch, winter 1852-1853. USNM 96 (one set of pharyngeal arches in osteological collection).
CAS 29301 (old IU 8628) (1) indicated as "type," but locality listed as Texas.
REMARKs.-Substitute name for Moniana nitida Girard 1857 (preoccupied in Notropis by
Alburnus nitidus Kirtland 1854). Types and type locality thus same as for Moniana nitida. Counts
for USNM 39657: pharyngeal teeth 4-4 in one, arches absent in other; anal ravs 7 in both. Counts
for USNM 96: pharyngeal teeth 1,4-4,0, with tooth in left lesser row represented by a stump.
Original description date 18 March 1896.
= Notropis braytoni Jordan and Evermann
Notropis brazosensis Hubbs and Bonham 1951
Hubbs and Bonham 1951:97-100, pl. 1 (fig. 2) and pl. 2 (fig. 2). TYPE LOCALITY: Brazos R., at
Wellborn Crossing, Brazos Co., Texas. HOLOTYPE: UMMZ 129827 (49.2), K. Bonham and party,
21 October 1938. PARATYPES: UMMZ 129828 (101), paratopotypes; UMMZ 129840 (18), UMMZ
146293 (2).
REMARKs.-Gilbert and Bailey (1962) synonymized this form with N. shumaordi. Original
description date 30 March 1951.
= Notropis shumardi (Girard)
BULLETIN FLORIDA STATE MUSEUM
Vol. 23, No. 1
Notropis brimleyi Bean 1903
Bean 1903:913-914. TYPE LOCALITY: Cane R., Cane River P.O. (near Burnsville), Yancey Co.,
North Carolina. HOLOTYPE: USNM 50601 (76.5), H. H. Brimley and F. Sherman, Jr., 4 October
1902. PARATYPES: None located. Original description listed two paratypes collected with holo-
type.
REMARKS.-Gilbert (1964:124) svnonvmized this form with N. coccogenis. Smith (1907:253)
erroneously gave the type locality as Cane Creek, in the French Broad River system, but Men-
hinick, Burton, and Bailey (1974:24) showed that the Cane River and Bollings (= Bowlens) Creek
mentioned in Bean's (1903) original description are in Yancey County, in the Nolichucky sys-
tem. Original description date 6 July 1903.
= Notropis coccogenis (Cope)
Leiciscus bIbalinuts Baird and Girard 1853
Baird and Girard 1853:391. TYPE LOCALITY: Otter Cr., trib. to N. Fk. of Red R., either Till-
man or Kiowa County, southwestern Oklahoma. (Locality erroneously listed in original descrip-
tion as in Arkansas). tHOLOTYPE: USNM 125 (one set of pharyngeal arches in bone collection),
R. B. Marcy and G. B. McClellan, 1852.
REMARKS.-Girard (1858:265) listed only one type specimen. Pharyngeal-tooth count for
USNM 125:4-4. Hubbs and Ortenburger (1929a:34) were first to synonymize this form with N.
lutrensis and, as first revisers, selected name lutrensis over bubalinus, both species having been
described on the same page of Baird and Girard's paper. Original description date 5 September
1853.
= Notropis lutrensis (Baird and Girard)
Notropis hairdi buccula Cross 1953
Cross 1953:252-259, fig. 1. TYPE LOCALITY: Brazos R., ca. 7 mi. S of Mineral Wells, at U.S.
hwy. 281 crossing, Palo Pinto Co., Texas. HOLOTYPE: KU 2642 (41.3), G. A. Moore and F. B. Cross,
14 April 1952. PARATYPES: KU 2318 (originally 13, now 9), UMMZ 166743 (ex KU 2318) (2),
USNM 163279 (ex KU 2318) (2), paratopotypes.
REMAHKS.-FoTrm very close to N. bairdi and perhaps not specifically distinct. Original descrip-
tion date 29 June 1953.
= Notropis buccula Cross
Notropis hbchanani Meek 1896
Meek 1896:342. TYPE LOCALITY: small creek near Poteau, Le Flore Co., Oklahoma. SYNTYPES:
USNM 47532 (3, 27.4-30.2) and USNM 125161 (3, 26.9-30.0) are catalogued as syntypes, but on
the basis of available evidence (see below) should not he so considered.
REMAHaKs.-Confusion attends the type material of N. hrchlanani. In the original description
Meek (1896:342) listed 14 syntypes from a small creek near Poteau, Oklahoma, and gave the
catalogue number as USNM 47532. At present six specimens are labelled "types" in the USNM
collection, three of which bear the above catalogue number and three of which are catalogued
as USNM 125161, the latter having been transferred from the old U.S. Bureau of Fisheries dupli-
cate collection. The labels in both jars say "Red River, Arthur, Texas," and nothing in the jars,
the catalogue, or the fil[ M,..L-,, these locality data to be in error.
The Red River collection was treated by Meek in the same paper as that from Poteau, and
N. buchanani is known to occur at both localities. Inquiries at the U.S. National Museum, the
Field Museum of Natural History, the California Academy of Sciences, and the University of
Michigan (those collections most likely to have Meek material) have failed to uncover any speci-
mens of N. buchanani from the Poteau locality in either their type or general collections. The
most likely explanation for this mixup is that Meek originally intended to use the Poteau speci-
mens as types, subsequently decided to use the Red River material instead (the latter specimens
are in excellent condition for their age), but for some reason failed to change the type locality
GILBERT: NOTROPIS TYPE CATALOGUE
in his paper. Whatever the reason, the Red River specimens cannot properly be regarded as
types, at least until such time as an explanation for the present situation is forthcoming.
Hubbs and Greene (1928; 377-378) reduced this form to a subspecies, but Bailey (1951:193)
subsequently re-elevated it to a full species. Original description date 13 April 1896.
= Notropis buchanani Meek
Notropis burchi Fowler 1942
Fowler 1942:4-6, figs. 5-6. TYPE LOCALITY: Mill Cr., trib. to St. Marvs R., Charlton Co.,
Georgia. HOLOTYPE: ANSP 69970 (47.5), J. W. Burch, 14 December 1940. PARATYPE: ANSP
69971 (1), paratopotype.
REMARKS.--Although Fowler designated a holotype, the above types are in the same jar and
are exactly the same length, with no indication which specimen is which. However, one speci-
men is in slightly better condition, and thus is here considered to be the holotype. Counts for
holotype: pharyngeal teeth 4-4; anal rays 8. Bailey, Winn, and Smith (1954:127) erroneously
synonymized this form with N. chalybaeus. Original description date 16 September 1942.
= Notropis imaculatus (Hay)
Notropis atherinoides caddonis Meek 1891
Meek 1891:136. TYPE LOCALITY: Little Red River, Judsonia, White County, Arkansas. TYPES:
None located.
REMARKs.-Other than the original description, the only reference to this form of which I
am aware is that by Meek (1894:78) (as N. atherinoides caddoensis). To my knowledge, it has
not previously been formally svnonymized with N. atherinoides. Original description date 19
June 1891.
= Notropis atherinoides Rafinesque
Photogenis caeruleus Jordan 1877
Jordan 1877c:338-339. TYPE LO( ALITY: tribes. of Oostanaula R. (primarily Rocky Cr.), above
Rome, Floyd Co., Georgia. LEC'TOTYPE: USNM 17883 (59.6), D. S. Jordan and C. H. Gilbert,
July 1876. Lectotype herein designated by R. H. Gibbs, Jr. LECTOPARATYPES: USNM 20114 (2),
MCZ 24396 (1), CU 1488 (1), BMNH 1880.1.21.10 (1), MNHN A.1306 (1), paratopotypes. The
three syntypes listed (one illustrated) by Fowler (1910:282; pl. 28, fig. 26) cannot be located on
the shelves at the ANSP, nor can a number for such specimens be found in the ANSP catalogue.
REMAKxs.-Gibbs (1955) reviewed species, but (1955:177) did not list CU, ANSP, nor BMNH
types, although Fowler's (1910: pl. 18, fig. 26) illustration of one of the ANSP syntypes (speci-
mens not seen by me) appears to be N. caenrleus. Name first appeared as a nomen nudum (Cy-
prinella caerulea) in Jordan and Copeland (1877:153). Original description date (30) June 1877.
= Notropis caeruleus (Jordan)
Notropis calientis Jordan and Snyder 1899
Jordan and Snyder 1899:122-123, fig. 4. TYPE LOCALITY: Rio Verde, Aguascalientes, Mexico.
HOLOTYPE: CAS-SU 6193 (40.2), J. O. Snyder, 9 January 1899. PARATYPES: CAS-SU 6198 (11),
BMNH 1900.9.29.152-157 (6), paratopotypes.
REMARKS.-Counts for holotype: Pharyngeal teeth 4-4; anal rays 7; body-circumferential
scales 12-2-11 = 25; caudal-peduncle circumferential scales 5-2-5 = 12. Counts for CAS-SU para-
types: anal rays 7 (10), 8(1). Original description date 30 August 1899.
= Notropis calientis Jordan and Snyder
Episema callisema Jordan 1877
Jordan 1877c:363-364. TYPE LOCALITY: S.Fk. of Ocmulgee R., Flat Rock, DeKalb Co., Geor-
gia. LECTOTYPE: USNM 17864 (55.4), D. S. Jordan and C. H. Gilbert, July 1876. Lectotype herein
designated by R. H. Gibbs, Jr. LECTOPARATYPES: USNM 163954 (ex USNM 17864) (3), USNM
20126 (1), MCZ 24384 (1), BMNH 1880.1.21.20 (1), paratopotypes.
BULLETIN FLORIDA STATE MUSEUM
REMARKS.-Gibbs (1955) reviewed species, but did not (1955:138) list MCZ nor BMNH types.
Name first appeared as a nomen nudum (Moniana callisema) in Jordan and Copeland (1877:154).
Original description date (30) June 1877.
= Notropis callisema (Jordan)
Photogenis callistius Jordan 1877
Jordan 1877c:337-338. TYPE LOCALITY: tribs. of Etowah and Oostanaula rivers, near Rome,
Georgia. (Most specimens from Silver Cr., trib. to Etowah R.) LECTOTYPE: MCZ 24363 (80.0),
D. S. Jordan and C. H. Gilbert, summer 1876. Lectotype herein designated by R. H. Gibbs, Jr.
LECTOPARATYPES: Possibly none (see below). One specimen each of BMNH 1880.1.21.67 and
MNHN A. 1289 were not examined by either Gibbs or me.
REMARKS.-Gibbs (1955) reviewed species. Of the seven extant former syntypes of this spe-
cies, two have not yet been examined (BMNH 1880.1.21.67 and MNHN A. 1289), two are N.
trichroistius (ANSP 19837-19838), two are N. caeruleus (USNM 17882 and 163965), one is Semo-
tilus atromaculatus (USNM), and only one is N. callistus as now recognized (MCZ 24363). Name
first appeared as a nomen nudum (Cyprinella callistia) in Jordan and Copeland (1877:153). Orig-
inal description date (30) June 1877.
= Notropis callistius (Jordan)
Notropis callitaenia Bailey and Gibbs 1956
Bailey and Gibbs 1956:1-14, fig. 1. TYPE LOCALITY: Flint R., ca. 1 mi. S of Radium Springs
outlet, 5.5 mi. S of Albany, Dougherty Co., Georgia. HOLOTYPE: UMMZ 168938 (64.4), H. E.
Winn and R. R. Rosanio, 20-29 September 1952. PARATYPES: UMMZ 163922 (70), USNM 171351
(ex UMMZ 163922) (6), ANSP 73864 (20), paratopotypes. Numerous other specimens examined,
but not designated paratypes.
REMARKS: Original description date 13 August 1956.
= Notropis callitaenia Bailey and Gibbs
Cyprinella calliura Jordan 1877
Jordan 1877e:61-62. TYPE LOCALITY: Alabama River system, Selma, Alabama. Type locality
given in original description as Black Warrior River, but this stream is some distance from Selma.
LECTOTYPE: USNM 6865 (89.6), Prof. Winchell. Gibbs (1957:192) designated lectotype. LECTO-
PARATYPEs: USNM 163951 (7), USNM 214661 (1), paratopotypes.
REMARKS.-Jordan (1885b:813) was first to synonymize this form with N. cercostigma stig-
maturus (= N. venustus stigmaturus).A jar labelled Cyprinella calliura, with appropriate locality
data and also bearing the catalogue number USNM 6865, was found in the non-type section of
the collection on 15 June 1975. This specimen, which I identify as N. venustus, presumably was
an original syntype that had, for reasons unknown, been placed in a separate jar some years ago.
Because it was not included among the lectoparatypes listed by Gibbs (1957:192), it was not
added to the series USNM 163951 but instead was recatalogued as USNM 214661. Original de-
scription date (31 December) 1877.
= Notropis venustus: cercostigma x stigmaturus
Cliola camura Jordan and Meek 1884
Jordan and Meek 1884:474-475. TYPE LOCALITY: Uncertain. According to the original descrip-
tion, the type locality is Fort Lyon, Colorado. This is well west of the present range of the spe-
cies, although N. camurus does occur in the middle and lower parts of the Arkansas River sys-
tem, and it is possible that a relict population was, till recently, present at the Fort Lyon locality.
A second possibility is Lyons, Kansas (also in the Arkansas system), as N. camurus occurs today
within about a hundred miles of there. The most likely possibility is that the types are from
Lyons County, in the Neosho River drainage (Arkansas R. system) of eastern Kansas, where the
species is common today. LECTOTYPE: USNM 15256 (80.0), Dr. E. Palmer. Gibbs (1961:345) desig-
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
nated lectotype. LECTOPARATYPES: None. Gibbs (1961:345) reidentified second syntype as Hybog-
nathus placitus.
REMARKs.-Gibbs (1961) reviewed species. Original description date 23 October 1884.
= Notropis camurus (Jordan and Meek)
Notropis cayuga Meek 1889
Meek 1889:305-307. TYPE LOCALITY: Cayuga Lake and Fall Creek, Ithaca, Tompkins Co.,
New York. SYNTYPES: CAS-SU 3790 (15, 39.3-46.5) (=N. bifrenatus), CAS-SU 67156 (36, 24.9-
50.2) (= N. heterodon), CAS-SU 67159 (1, 35.2) (= N. analostanus), S. E. Meek, 1885-1886. Speci-
mens were identified by me and subsequently recatalogued by species, but all are still syntypes.
REMARKS.-The above syntypes were originally in two lots, both apparently with the same
data, which were mixed as a result of the 1906 earthquake. Hubbs (1926:41) restricted the species
name cayuga to N. bifrenatus, so that the lectotype, when selected, should come from CAS-SU
3790. Original description date (31 December) 1889.
= Notropis bifrenatus (Cope)
Notropis celayensis Alvarez 1958
See page 22.
Hypsilepis cornutus cerasinls Cope 1868
Cope 1868a:159. TYPE LOCALITY: headwaters of Roanoke R., Virginia. LECTOTYPE: ANSP
3833 (73.0), E. D. Cope. Gilbert (1964:137) designated lectotype. Fowler's (1910) earlier lecto-
type designation (ANSP 3791) is discussed below. LECTOPARATYPES: ANSP 3792-3799 (8), parato-
potypes. Remainder of original syntypic series (ANSP 3791, 3800-3833) (34 specimens) reidenti-
fied (Gilbert 1964:137) as N. albeolus and catalogued under that name.
REMARKs.-Gilbert (1964:137-140) reviewed species. Fowler (1910: pl. 18, fig. 25) illustrated
the "type" of Hypsilepis cornutus cerasinus which, according to its calculated length (99 mm SL),
obviously was based on one of the specimens of N. albeolus in the original syntypic series. Should
Fowler's lectotype designation be allowed to stand, it could result in the changing of two long-
established species' names. In the interest of nomenclatural stability, I (Gilbert 1977) therefore
have requested that the International Commission suspend Fowler's (1910) designation and allow
mine (Gilbert 1964) to stand. Inasmuch as the same catalogue number (ANSP 3791) was used for
two different lectotype specimens, a new catalogue number (ANSP 3833) was required for the
lectotype I designated in 1964. Original description date 11 May 1868.
= Notropis cerasinus (Cope)
Cyprinellh cercostigma Cope 1868
Cope 1868a:157. TYPE LOCALITY: Pearl R., Monticello, Lawrence Co., Mississippi. TYPES:
None located. Four specimens supposedly at USNM, collected by Helen Tennison.
REMARKs.-Gibbs (1957:189-192) reviewed this form. In the original description Cope gave
the pharyngeal tooth count as 2,4-4,2, which if accurate would refer to a species other than N.
cenustus. Original description date 11 May 1868.
= Notropis venustus cercostigmo (Cope)
Hybopsis chalybaeus Cope 1869
Cope 1869:383-384. TYPE LOCALITY: trib. of Schuylkill R., near Conshohocken, Montgomery
Co., Pennsylvania. TYPES: None located. Fowler (1909, 1910) did not list types in ANSP collec-
tion.
REMARKs: Swift (1970) reviewed species. Original description date (31) August 1869.
= Notropis chalybaeus (Cope)
Notropis chamberltini Evermann 1898
Evermann, in Jordan and Evermann 1898:2800-2801. TYPE LOCALITY: Atchafalaya R., Mel-
BULLETIN FLORIDA STATE MUSEUM
ville, St. Landry Psh., Louisiana. HOLOTYPE: USNM 48900 (70.0), F. M. Chamberlain, 5 May
1897. PARATYPES: USNM 48901(4), USNM 125608(5), CAS-SU 8570(4), paratopotypes.
REMARKs.-Ortenburger and Hubbs (1927:126) were first to synonymize this form with N.
illecebrosus (= N. shumardi). Original description date (30) November 1898.
= Notropis shumardi (Girard)
Luxilus chickasavensis Hay 1881
Hay 1881:506. TYPE LOCALITY: Chickasawha R., Enterprise, Clark Co., Mississippi. LECTO-
TYPE: USNM 27419 (76.6), O. P. Hay, March-April 1880. Gibbs (1957:190) designated lecto-
type. LECTOPARATYPES: USNM 163953 (8), paratopotypes.
REMARKS.-Gibbs (1957:190) synonymized this form with N. cenustus cercostigmla. Original
description date 23 February 1881.
= Notropis venustus cercostigma (Cope)
Notropis chihuahua Woolman 1892
Woolman 1892:260. TYPE LOCALITY: Rio de los Conchos, Chihuahua City, Chihuahua, Mex-
ico. SYNTYPES: UMMZ 61702 (ex old IU 4868) (5,46.7-59.0), CAS 14213 (ex old IU 4868) (19, 43.2-
56.1), CAS-SU 818 (27, 41.0-55.4), USNM 44151 (10, 43.6-56.3), USNM 125186 (1, 49.6), FMNH
1889 (1, 53.7), BMNH 1894.1.27.48-57 (12, 40.0-51.0), A. J. Woolman and J. T. Scovell, July-
August 1891.
REMARKS: Original description date (31) March 1892.
= Notropis chihuahua Woolman
Hybopsis chiliticus Cope 1870
Cope 1870:462. TYPE LOCALITY: trib. of Yadkin R., Roane (= Rowan) Co., North Carolina.
LECTOTYPE: ANSP 4378 (49.0) E. D. Cope, Fall 1869. Fowler (1910: pl. 19, fig. 36) designated
lectotype by illustrating "type" (ANSP 4378); calculated length 49.5 mm SL. LECTOPARATYPES:
ANSP 43794388 (10), paratopotypes.
REMARKS.-Condition of types fair. Counts for syntypes: pharyngeal teeth 1,4-4,1 (6), 2,4-4,1
(1), 1,4-4,2 (1), 2,44,2 (3); anal rays 8 (9), 9 (2). Original description date 21 November 1870.
= Notropis chiliticus (Cope)
Cliola chlora Jordan 1878
Jordan 1878c:791. TYPE LOCALITY: Upper Missouri River system. (Exact locality unknown;
see opening remarks by Jordan [1878c:777]). SYNTYPES: USNM 20193 (9, 36.843.4) (in two jars:
eight specimens in first jar, one specimen in second), Elliott Coues, 1873 or 1874.
REMARKS.-Pharyngeal teeth 4-4 in two, -4 in two; anal rays 7 in all nine. Jordan and Ever-
mann (1896:263) synonymized this form with N. scylla, and Hubbs and Greene (1928:375) synony-
mized both with the Great Plains form (missuriensis) of N. delicious. Original description date
11 December 1878.
= Notropis stramineus missuriensis (Cope)
Codoma chloristia Jordan and Brayton 1878
Jordan and Brayton 1878:21-23. TYPE LOCALITY: Saluda R., Farr's Mills, W of Greenville,
Greenville Co., South Carolina. LECTOTYPE: MCZ 24380 (50.1), D. S. Jordan and C. H. Gilbert,
summer 1877. Gibbs (1963:522) designated lectotype. LECTOPARATYPE: BMNH 1880.1.21.43 (1),
paratopotype.
REMARKS.-Gibbs (1963:522-524) reviewed this form and regarded it as a subspecies of N.
analostanus, but most still consider it to he a valid species (Bailey et al. 1970:21). Original descrip-
tion date (31 December) 1878.
= Notropis chloristius (Jordan and Brayton)
Hybopsis chlorocephalus Cope 1870
Cope 1870:461. TYPE LOCALITY: tribs. of Catawba R., North Carolina. LECTOTYPE: ANSP
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
2755 (44.0), E. D. Cope, fall 1869. Fowler (1910: pl. fig. 35) designated lectotype by illustrating
"type" (ANSP 2755); calculated length 45.1 mm SL. LECTOPARATYPES: ANSP 2756-2767 (12),
ANSP 2803-2829 (27). One original syntype (ANSP 2768) reidentified as Hybopsis hypsinotus and
recatalogued under that name.
REMARKS: Condition of types fair (ANSP 2755-2767) to poor (ANSP 2803-2829), the bodies
soft in the latter series. Counts for lectotype: pharyngeal teeth 1,4-; anal rays 8. Counts for ANSP
2756-2767: anal rays 8 (11), 9 (1). Counts for ANSP 2803-2829: teeth 1,4-4,1 (5), 1,4-4,0 (1); anal
rays 8 (27). Original description date 21 November 1870.
= Notropis chlorocephalus (Cope)
Notropis altipinnis chowcanus Hubbs and Ranev 1948
Hubbs and Raney 1948:10-11. TYPE LOCALITY: Waqua Cr., trib. to Nottoway R., near Raw-
lings, Brunswick Co., Virginia. HOLOTYPE: UMMZ 94515 (38.2), Donald Ameel, 10 November
1931. PARATYPES: UMMZ 94516 (5), USNM 101338 (9), paratopotypes; CU 10569 (75 originally,
now 65).
REMARKs.-This subspecies and others described in the same paper are not usually regarded
as valid, although they have never been formally synonymized. Original description date 26
April 1948.
= Notropis altipinnis (Cope)
Hyhopsis chrosonmus Jordan 1877
Jordan 1877c:333-334. TYPE LOCALITY: tribes. of Etowah and Oostanaula rivers, around Rome,
Georgia. (Most specimens from Silver Cr., trib. to Etowah R.). LECTOTYPE: USNM 17881 (44.5),
D. S. Jordan and C. H. Gilbert, July 1876. Lectotype herein designated by C. C. Swift. LECTO-
PARATYPES: USNM 203857 (ex USNM 17881) (1), ANSP 19843-19845 (3), CU 1487 (1), MCZ
24373 (1), BMNH 1880.1.21.40 (1), MNHN A. 1292 (3), paratopotypes.
REMARKs.-Swift (1970) reviewed species. Fowler (1910: pl. 20, fig. 41) illustrated "cotype."
Name first appeared as a nomen nuduim (Hybopsis chros(mugs) in Jordan and Copeland
(1877:151). Original description date (30) June 1877.
= Notropis chrosomius (Jordan)
Luxilus chrysoccphalus Rafinesque 1820
Rafinesque 1820b:48. TYPE LOCALITY: "Kentucky." Neotype locality is a creek ca 6 mi. SSW
of Danville, off st. hwy. 35, Lincoln Co., Kentucky. NEOTYPE: UMMZ 174539(99.0), R. M.
Bailey and party, 5 April 1953. Gilbert (1964:160) designated neotype.
REMARKS.-Jordan (1876a:94) was first to synonymize this form with N. cornutus. Hubbs
(1926:46) regarded it as a subspecies of N. cornutus, and Gilbert (1961a, 1961b) elevated it to a
full species. Menzel (1970, 1976), on the basis of close similarity to N. cornutus in blood serum
protein pattern, recommended reversion to status of a subspecies of N. cornutus. Gilbert
(1964:157-167) reviewed species. Original description date (31) May 1820.
= Notropis chrysocephalus chrysorephalus (Rafinesque)
Leuwiscus coccogenis Cope 1868
Cope, in Giinther, 1868:253. TYPE LOCALITY: Holston R., Virginia. LECTOTYPE: USNM 36849
(97.3), E. D. Cope, September 1867. Gilbert (1964:123) designated lectotype. LECTOPARATYPE:
MCZ 35653 (1), BMNH 1868.1.10.3 (8), ANSP 5379 (1), ANSP 3561-3650 (90), MNHN 4852 (1)
paratopotypes. Remainder of larger ANSP syntypic series (ANSP 3651-3660) comprises four
other species (N. chrysocephalus, N. galacturus, N. rubricroceus, and N. telescopes), which are
now recatalogued under those names.
REMARKS.-Gilbert (1964:122-126) reviewed species. Original description date 14 March
1868.
= Notropis coccogenis (Cope)
1978
BULLETIN FLORIDA STATE MUSEUM
Notropis cummingsae collis Hubbs and Raney 1951
Hubbs and Raney 1951:16-22, pl. 1, fig. 3. TYPE LOCALITY: Roses Cr., 1.5 mi. S of Smyrna,
on st. hwy. 181, Burke Co., North Carolina. HOLOTYPE: UMMZ 160557 (34.0), E. E. Brown, 6
September 1946. PARATYPES: UMMZ 160558 (2), CU 11218 (3), paratopotypes; UMMZ 94553
(1), UMMZ 94556-94557 (18), CU 11269 (9).
REMARKS.-This is a weakly defined form, differing from the nominate subspecies in usually
having one less anal ray. It is sometimes recognized, although the usual practice is to treat all
populations of N. cummingsae binomially. Original description date 11 December 1951.
= Notropis cummingsae Myers
Notropis comalis Jordan and Gilbert 1885
See page 22.
Moniana complanata Girard 1857
Girard 1857:200. TYPE LOCALITY: Rio Grande (or tributary), Brownsville, Cameron County,
Texas. SYNTYPE: USNM 94 (one pharyngeal arch), Capt. Van Vliet.
REMARKs.-Girard (1859a:56; pl. 31, figs. 17-20) listed three original syntypes and illustrated
one of them. Count for pharyngeal arch: 4-. Jordan (1885a:125) synonymized this form with N.
leoninus, and Jordan and Evermann (1896:272) later synonymized both with N. lutrensis. Con-
treras (1975) regarded it as a junior synonym of N. I. lutrensis. Original description date 25 April
1857.
= Notropis lutrensis lutrensis (Baird and Girard)
Rutilus compressus Rafinesque 1820
Rafinesque 1820b:51. TYPE LOCALITY: "Monongahela River." TYPES: None located.
REMARKS.-Possibly a synonym of N. cornutus or N. photogenis (see Jordan, Evermann, and
Clark 1930:123). Of these, N. cornutus is the more likely possibility, based on a compressed body
and dorsal and anal fin each with 9 rays, although Gilbert (1964:140-142) did not list it under
synonymy of that species. Original description date (31) May 1820.
= Not definitely identifiable. Most likely Notropis cornutus (Mitchill)
Notropis cooglei Hildebrand and Towers 1928
Hildebrand and Towers 1928:18. TYPE LOCALITY: Pelucia Cr., S of Greenwood, Leflore Co.,
Mississippi. HOLOTYPE: USNM 88379 (58.5), 19 June 1925, I. L. Towers. PARATYPES: USNM
127019 (38 originally, now 30), UMMZ 167162 (ex USNM 127019) (8), USNM 127021 (54 orig-
inally, now 46), UMMZ 167164 (ex USNM 127021) (8), USNM 127024 (26 originally, now 22),
UMMZ 167165 (ex USNM 127024) (4), paratopotypes; USNM 127020 (1), USNM 127022 (30
originally, now 26), UMMZ 167166 (ex USNM 127022) (4), USNM 127023 (72 originally, now
62), UMMZ 167163 (ex USNM 127023) (10).
REMARKS.-Bailey, Winn, and Smith (1954:128) synonymized this form with N. renustuis,
and Gibbs (1957:180) later synonymized it with N. v. venustus. Original description date 2 Febru-
ary 1928.
= Notropis cenustus cenustus (Girard)
Leuciscus copii Ginther 1868 emendedd spelling Lewiscus copei)
Ginther 1868:255-256. TYPE LOCALITY: Walker Cr., either Bland or Giles Co., Virginia; St.
Joseph R. and Dowagiac Cr. (trib. to St. Joseph R.), Berrien Co., Michigan. SYNTYPES: BMNH
1868.1.10.24 (3, 43.0-58.0) (Walker Cr.), E. D. Cope, August-September 1867; ANSP 2577 (1,
42.0), MCZ 4688 (2,50.6-55.2) (St. Joseph R. and Dowagiac Cr.), E. D. Cope.
REMARKS: Giinther (1868:255-256) introduced the name copii to replace the species name
jaculus (preoccupied in Alburnellus) which, contrary to Giinther's indication, had not yet been
described (see Gilbert 1971a:476). Both names thus emanate from the same publication. Accord-
ing to W. I. Follett, whose advice I sought in the matter, Ginther should be considered the first
reviser, as he chose the name copii over jaculus and clearly indicated that they represent the same
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
species. (This has been confirmed by re-examination of the types.) Alburnellus jaculus thus stands
as a junior synonym of Leuciscus copii. The name jaculus, though first appearing as a junior syn-
onym, is available under the rules of zoological nomenclature (1964: Article 11D), as a result of
Cope's (1869:387) subsequent description. Although the Michigan localities were not specifically
mentioned in the original description, they are still regarded as type localities and the specimens
taken there as syntypes, for reasons discussed earlier in this paper, and because the range of the
species was specified in the original description to include Michigan and Virginia. Original de-
scription date 14 March 1868.
= Notropis ruhellus (Agassiz)
Cyprinus cornutus Mitchill 1817
Mitchill 1817:289. TYPE LOCALITY: Walkill R., New York. Neotype locality is Walkill R.,
3 mi. SW of New Paltz, Ulster Co., New York. NEOTYPE: UMMZ 174540 (69.0), J. R. Greeley and
R. M. Bailey, 2 June 1936. Gilbert (1964:142) designated neotype.
REMARKS.-Gilbert (1964:140-151) reviewed species. See subsequent account of Cyprinus
megalops for more detailed discussion of nomenclature of C. comutus. On the basis of close simi-
larity in blood serum protein pattern between this form and N. chrysocephalus, Menzel (1970,
1976) recommended calling the latter a subspecies of N. cornutus. Original description date (31)
August 1817.
= Notropis cornutus (Mitchill)
Moniana couchi Girard 1857
Girard 1857:201. TYPE LOCALITY: Rio San Juan, vicinity of China, Nuevo Leon, Mexico.
SYNTYPES: USNM 103 (= USNM 20227 ?) (33, 18.7-42.8 + one set of pharyngeal arches in bone
collection and one unidentified specimen not this species), MCZ 1795 (2, 31.0-39.6), Lt. D. N.
Couch, 1853. ANSP 2924 (1, 39.2) is also labelled as a type of this species, but label in jar says
"Arkansas River near Fort Smith. Dr. George B. Shumard. Smiths. Inst. 2982." This obviously
should not be considered a syntype of this species.
REMARKS.-Girard (1859a:57; pl. 30, figs. 21-24) listed 20 original syntypes and illustrated
one of them. Jordan (1885a:125) was first to synonymize this form with N. lutrensis. Contreras
(1975) regarded it as a junior synonym of N. 1. lutrensis. USNM 103 erroneously recatalogued
as USNM 20227 at one time. Counts for USNM syntypes: teeth 4-4 in two; anal rays 9(3), 10(1).
Counts for ANSP syntypes: teeth 4-4 and anal rays 9 in both. Original description date 25 April
1857.
= Notropis lutrensis lutrensis (Baird and Girard)
Leuciscus croceus Storer 1845
Storer 1845:48. TYPE LOCALITY: "Alabama." TYPES: None located.
REMARKS.-Possibly not a Notropis, although dorsal-ray count (8), anal-ray count (7), and
orange fins suggest N. longirostris. Original description date (31 December) 1845.
= Not definitely identifiable, but most likely Notropis longirostris (Hay)
Ceratichthys cumingii Giinther 1868 emendedd spelling Ceratichthys cumingi)
Giinther 1868:177-178. TYPE LOCALITY: Unknown. Label in jar says "California," which is
obviously in error, as N. boucardi does not occur in California. SYNTYPES: BMNH 1856.4.9.10
(2, 50.0-57.5), H. Cuming.
REMARKS.-Specimens identified as Hybopsis houcardi by C. L. Hubbs on 30 September 1958.
The species cumingii and boucardi were described in the same publication, and I, as first re-
viser, hereby select the latter name. Original description date 14 March 1868.
= Notropis boucardi (Giinther)
Notropis cummingsi Myers 1925
Myers 1925:14, fig. 1. TYPE LOCALITY: Upper Burnt Mill Cr., Wilmington, New Hanover
County, North Carolina. HOLOTYPE: AMNH 8391 (48.4), G. S. Myers, 19 May 1924. This speci-
BULLETIN FLORIDA STATE MUSEUM
men cannot be located and may be lost (C. L. Smith, in litt., 23 November 1971). PARATYPES:
USNM 94242 (1), ANSP 53943 (1), UMMZ 66514 (4), CAS-SU 24327 (1), BMNH 1925.10.19.5 (1),
paratopotypes.
REMARKs.-Hubbs and Raney (1951) reviewed this species and recognized two subspecies
(collis and the nominate form). Although these have sometimes been considered valid (Smith-
Vaniz 1968: fig. 87), the usual practice is to treat all populations of N. cummingsae binomially.
Hubbs and Raney (1951:5) also emended the name ending to conform to the International Com-
mission on Zoological Nomenclature ruling, as species was named for Mrs. J. H. Cummings.
Original description date 23 April 1925.
= Notropis cummingsae Myers
Hypsilepis cornutus cyaneus Cope 1868
Cope 1868a:160. TYPE LOCALITY: Montreal R., (trib. to Lake Superior), Keeweenaw Pt.,
Keeweenaw Co., Michigan. LECTOTYPE: ANSP 3950 (119.0), Dr. J. H. Slack. Fowler (1910: pl. 18,
fig. 32) designated lectotype by illustrating "type" (ANSP 3950); calculated length 124.0 mm SL.
LECTOPARATYPES: ANSP 3951-3959 (9), paratopotypes.
REMARKS.-Hubbs (1926:46) synonymized this form with N. cornutus frontalis (= N. cornu-
tus). Condition of syntypes good. Original description date 11 May 1868.
= Notropis corutus (Mitchill)
Lythrurus cyanocephalus Copeland 1877
Copeland, in Jordan, 1877a:70-71. TYPE LOCALITY: Root R., Racine Co., Wisconsin. (See
under Remarks). LECTOTYPE: USNM 17857 (45.0), Dr. Hoy, 1874. Snelson and Pflieger (1975:235)
designated lectotype.
REMARKs.-Snelson and Pflieger (1975) reviewed this form. Name first appeared as a nomen
nudum (Lythrurus cyanocephalus) in Nelson (1876:47). The original type specimens came from
two localities: the one listed above, and Bass Creek, tributary to the Rock River, near Hanover,
Wisconsin (coll. H. E. Copeland). Inasmuch as only one syntype (USNM 17857) apparently re-
mains, it has been chosen as the lectotype and the place from which it came (Root River) the type
locality. Original description date 17 April 1877.
= Notropis umbratilis cyanocephalus (Copeland)
Moniana deliciosa Girard 1857
Girard 1857:199. TYPE LOCALITY: Leon R. (= Leon Cr.), trib. to San Antonio R., Bexar Co.,
Texas. LECTOTYPE: USNM 119 (49.6), Dr. C. B. Kennerly, 1853. Clark Hubbs (1954a:73) desig-
nated lectotype. LECTOPARATYPES: None. Suttkus (1958:310) reidentified, as N. colucellus noco-
mis, the specimen (USNM 162723) Clark Hubbs (1954a:73) listed as a paratype.
REMARKS.-Girard (1858:275) listed only 12 original syntypes. Clark Hubbs (1954a:72-73) be-
lieved the lectotype and lectoparatype of Moniana deliciosa to be the species now known as N.
stramineus (the fish to which the name deliciostis was applied for many years). Nine other syn-
types were recatalogued as N. colucellus (USNM 162724). Suttkus (1958:307-318) later reidenti-
fied the lectotype as N. texanus, the lectoparatype as N. volucellus nocomis, and one of the nine
recatalogued syntypes as N. texanus (the other eight syntypes are N. colucellus, as indicated by
Hubbs). Suttkus also identified the three other extant syntypes as N. volucellus nocomis: MCZ
1690 (2) and ANSP 4170 (1). Thus, none of the extant syntypes are N. stramineus. Counts for
lectotype: pharyngeal teeth 2,44,2; anal rays 7. Fowler (1910: pl. 15, fig. 4) illustrated ANSP
"cotype." Original description date 25 April 1857.
= Notropis texanus (Girard)
Alburnus dilectus Girard 1857
Girard 1857:193. TYPE LOCALITY: Arkansas R., near Ft. Smith, Arkansas. SYNTYPES: USNM
71 (2, 56.8-63.8), USNM 36927 (27, 39.0-60.2 + two specimens of Notropis colucellus), MCZ
1785 (2, 51.5-52.0), Dr. George C. Shumard, 14 July 1853.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
REMARKS.-Girard (1858:259; pl. 57, figs. 9-12) listed only 24 original syntypes and illustrated
one of these. This form was first svnonymized with N. atherinoides by Jordan (1878a:422), hut
was subsequently regarded as a distinct species, closely related to N. nibrifrons (= N. rubellus)
(Jordan and Evermann 1896:294). Hubbs and Ortenburger (1929b:83) again svnonymized it with
N. atherinoides. Knapp (1953:61) considered it to be a subspecies of N. atherinoides. Original
description date 25 April 1857.
= Notropis atherinoides Rafinesque
Minnilus dinenmus Rafinesque 1820
Rafinesque 1820b:46. TYPE LOCALITY: "Ohio River." TYPES: None located.
REMARKS.-Jordan and Copeland (1877:154) synonymized this form with Alburnellus jaculus,
and Jordan (1878a:422) synonymized it (dinemuis) with N. atherinoides. Jordan and Evermann
(1896:293) later synonymized both with N. atherinoides. Original description date (31) May 1820.
= Notropis atherinoides Rafinesque
Hypsilepis diplaemia Cope 1868
Cope 1868a:162-163. TYPE LOCALITY: Grand River (or tributary thereof), Lansing, Ingham
Co., Michigan. SYNTYPES: Possibly ANSP 3054-3055 (2, 49.0-50.0). These specimens hear only
the data "Michigan, E. D. Cope," with no positive indication that they actually are types.
REMARKs.--Strictly speaking, Rafinesque (1820b:50) was first to use this specific name in his
description of Senotilus diplemia, a form that cannot be positively identified, but which prob-
ably does not refer to a species of Notropis. Others (Kirtland 1846:276, pl. 22, fig. 3; Putnam
1863:7; Gunther 1868:250) subsequently applied it to a variety of species, and Cope (1868a:162-
163) also regarded Rafinesque as authority for the name. Jordan (in Jordan and Meek 1884:476),
however, proposed the substitute name lythnrurs for Hypsilepis diplaemia Cope, at the same time
clearly indicating this was not the Semotilus diplemiia of Rafinesque. According to Article 16a,
International Code of Zoological Nomenclature, Jordan's action allows Cope's (1868a) descrip-
tion of H. dipkiemia to stand as the original description of N. lythrnrus, which in turn is a junior
synonym of N. nmbratilis cyanocephalus (Copeland 1877). Although Jordan and Gilbert (1877:95)
designated Semotilus diplemia Rafinesque 1820 as type species of the genus Lythnrurs, the spe-
cies involved actually is Hypsilepis diplaemia Cope 1868, for reasons discussed above. For further
discussion see accounts of t. lythrurns (this paper), and also that by Snelson and Pflieger
(1975:235). Original description date 11 May 1868.
= Notropis umbratilis cyanocephalus (Copeland)
Hybopsis dorsalis Agassiz 1854
Agassiz 1854:358. TYPE LOCALITY: Burlington, Iowa. SYNTYPES: MCZ 1947 (93, 22.8-47.0),
USNM 120418 (ex MCZ 1947) (4,30.0-38.5) UMMZ 86506 (ex MCZ 1947) (10,30.0-48.0), Dr. I. H.
Ranch.
REMARKS.-This form was generally ignored after its original description, or was placed in
the questionable synonymy of N. delicious (=N. stramineus). Hubbs and Greene (1928:380)
were first to clarify its status. Types soft, but otherwise in good condition for old specimens.
Counts for two of MCZ syntypes: pharyngeal teeth 1,4-4,1 in both; anal rays 8 in both. Counts
for four USNM syntypes: pharyngeal teeth 1,4-4,1 (2); anal rays 7 (1), 8 (3). For remarks on sub-
species of N. dorsalis see account of Photogenis piptolepis. Original description date (31 Decem-
ber) 1854.
= Notropis dorsalis (Agassiz)
Notropis edwardraneyi Suttkus and Clemmer 1968
Suttkus and Clemmer 1968:18-39, figs. 1-10. TYPE LOCALITY: Alabama R. at Yellow Jacket
Bar, river mile 129.8, 1.2 mi down-river from Holly Ferry crossing, or 12.5 mi E of Pine Hill,
Wilcox Co., Alabama. HOLOTYPE: TU 49485 (57.6), R. D. Suttkus and G. R. Gunning, 8 March
1967. PARATYPES: USNM 202435 (100)), UMMZ 187475 (1(X)), ANSP 109424 (100), CU 52941
1978
BULLETIN FLORIDA STATE MUSEUM
(100), MCZ 45878 (100), CAS-SU 66551 (100), KU 12674 (100), UAIC 2791 (100), FMNH 74294
(100), TU 4618 (33), TU 40303 (1020), TU 40925 (85), TU 41400 (390), TU 41726 (16), TU 42737
(5630), TU 44028 (7324), TU 46802 (223), TU 47361 (49), TU 47924 (105), paratopotypes; TU
33381 (1613), TU 35243 (236), TU 35269 (314), TU 35323 (28), TU 40293 (297), TU 40320 (491),
TU 40335 (243), TU 40900 (289), TU 40940 (226), TU 40950 (109), TU 41608 (284), TU 41632
(92), TU 41639 (187), TU 41655 (119), TU 41666 (147), TU 41670 (301), TU 41695 (83), TU 41711
(29), TU 41732 (19), TU 41745 (63), TU 41755 (12), TU 41761 (9), TU 41772 (151), TU 41791 (22),
TU 41797 (4), TU 41813 (10), TU 41823 (1), TU 42731 (370), TU 42746 (687), TU 42759 (900),
TU 44011 (175), TU 44029 (94), TU 44045 (461), TU 46783 (202), TU 46796 (395), TU 46819 (43),
TU 46830 (32), TU 47346 (64), TU 47374 (86), TU 47387 (168), TU 47396 (33), TU 47408 (475),
TU 47420 (173), TU 47435 (229), TU 47452 (134), TU 47477 (89), TU 47491 (24), TU 47499 (16),
TU 47515 (306), TU 47762 (143), TU 47781 (9), TU 47822 (85), TU 47838 (75), TU 47909 (152),
TU 47936 (39), TU 47968 (23), TU 47980 (180), TU 47994 (65), TU 48012 (232).
REMARKs.-Original description date 16 October 1968.
= Notropis edwardraneyi Suttkus and Clemmer
Opsopoeodus emiliae Hay 1881
Hay 1881:507-508. TYPE LOCALITY: The original description listed three localities, all in
Mississippi: Catawha Cr., Artesia, Lowndes Co.; Chickasawha R. and tributaries, Enterprise,
Clarke Co.; and Horsehunter Cr., near confluence with Noxubee R., Macon, Noxubee Co. Gilbert
and Bailey (1972:20) restricted type locality to Horsehunter Creek (see REMARKS). SYNTYPES:
None located. Hay, in original description, listed one specimen from Artesia, one from Enter-
prise, and several from Macon. He listed USNM 27429 as type number, whereas Jordan and Ever-
mann (1896:248) gave it as USNM 32222. Presumably these numbers apply to different syntypic
series.
REMARKs.-Gilbert and Bailey (1972) reviewed N. emiliae, and reduced Opsopoeodus to a
subgenus of Notropis, an action questioned by Campos and Hubbs (1973) on the basis of differ-
ence in chromosome number. Inasmuch as the syntypes of N. emniliae from Macon were the ones
catalogued as USNM 27429 (Gilbert and Bailey 1972:20), they restricted the type locality on that
basis. Original description date 23 February 1881.
= Notropis emiliae emiliae (Hay)
Photogenis leucops engraulinus Cope 1868
Cope 1868a:164. TYPE LOCALITY: trib. to Kanawha R., Austinville, Wythe Co., Virginia.
HOLOTYPE: ANSP 3033 (length not determinable), E. D. Cope, August-September 1867.
REMARKS.-Gilbert (1971a:477) discussed the type specimen. Type in very poor condition
(head missing); anal rays 10, pelvic rays 9-9. Original description date 11 May 1868.
= Notropis photogenic (Cope)
Hudsonius euryopa Bean 1880
Bean 1880:285-286. TYPE LOCALITY: McBean Cr., near McBean, at Burke and Richmond
Co. line, Georgia. LECTOTYPE: USNM 23513 (85.2), A. Graves, 1879. Lectotype herein desig-
nated by William Seaman, Jr. The other syntype is N. petersoni and is recatalogued under that
name.
REMARKS.-Original description date 30 March 1880.
= Notropis hudsonius (Clinton)
Photogenis eurystomus Jordan 1877
Jordan 1877c:356. TYPE LOCALITY: Nancys Cr., tribe. to Chattahoochee R., Atlanta, Fulton
Co., Georgia. LECTOTYPE: MCZ 24388 (68.3), D. S. Jordan and C. H. Gilbert, summer 1876. Gibbs
(1957:190) designated lectotype. This specimen apparently is the only type still extant.
REMARKs.-Name first appeared as a notmen nudum (Cyprinella eurystoma) in Jordan and
Copeland (1877:153). Bailey, Winn, and Smith (1954:128) were first to synonymize this form
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
with N. venustus, and Gibbs (1957:189-190) svnonvmized it with N. renustus cercostigma. Con-
dition of type fair to good (wrinkled); anal rays 9. Original description date (30) June 1877.
= Notropis renustus cercostigma (Cope)
Notropis euryzonus Suttkus 1955
Suttkus 1955:85-100, fig. 1 (two illustrations). TYPE LOCALITY: Uchee Cr., 0.7 mi. E of Mar-
vyn, Lee Co., Alabama. HOLOTYPE: CU 28346 (49.0), R. D. Suttkus, R. H. Gibbs, Jr., and C. F.
Cole, 12 June 1949. PARATYPES: CU 15990 (36), paratopotypes; CU 1398:3 (5), CU 14316 (43),
CU 16194 (2), UMMZ 123951 (1), UMMZ 128744 (6), UMMZ 128745(1), TU 10700 (8), TU 10718
(60).
REMARKS.-Original description date 28 December 1955.
= Notropis euryzonus Suttkus
Notropis umhbratilis fasciolaris Gilbert 1891
Gilbert 1891:148. TYPE LOCALITY: Spring and Pin-hook creeks, Huntsville, Alabama. LEC-
TOTYPE: USNM 42504 (59.5), P. H. Kirsch, 27-28 May 1889. Lectotype herein designated by
F. F. Snelson, Jr. LECTOPARATYPES: USNM 203331 (ex USNM 42504) (4), USNM 43529(5), para-
topotypes; USNM 36185(7), USNM 36831(2), USNM 43485(11), CAS-SU 937(9).
REMARKs.-Smith-Vaniz (1968:42) was first to synonymize this form with N. ardens. Original
description date 16 June 1891.
= Notropis gardens (Cope)
Hudsonius fluciatilis Girard 1857
Girard 1857:210. TYPE LOCALITY: Chicago Harbor, Lake Michigan, Illinois; also Root River,
at Racine, Wisconsin. SYNTYPES: USNM 14 (2, 92.5-93.7 + one set of pharyngeal arches in bone
collection) (Chicago Harbor), S. F. Baird; USNM 13 (one set of pharyngeal arches in bone collec-
tion) (Root River), S. F. Baird.
REMARKS.-Counts for USNM 13: teeth 1,4-4,2. Counts for USNM 14; teeth 2,4-4,1 (arches in
bone collection). Jordan (1885a:127) was first to synonymize this form with N. hudsoniuls. Origi-
nal description date 25 April 1857.
= Notropis hudsonius (Clinton)
Cyprinella forbesi Jordan 1878
Jordan 1878a:57-58. TYPE LOCALITY: Illinois R., Union Co., Illinois. SYNTYPES: USNM 29869
(1, 50.8), CAS-SU 1290 (2, 53.2-57.4), S. A. Forbes, summer 1877.
REMARKS.-Jordan (1885b:812) was first to synonymize this form with N. lutrensis. Hubbs,
in Hubbs and Ortenburger (1929b:75) called it the "northern subspecies of N. lutrensis." Counts
for USNM 29869: teeth 1,4-4,0: anal rays 9. Counts for CAS-SU 1290: teeth 1,4-4,1 and 4-4; anal
rays 9 and 8, respectively. Original description date (30) June 1878.
= Notropis lutrensis (Baird and Girard)
Notropis forlonensis Meek 1904
Meek 1904:70-71, fig. 20. TYPE LOCALITY: Rio Forlon, tribe. to Rio Panuco, Forlon, Tamau-
lipas, Mexico. HOLOTYPE: FMNH 4478 (40.8), S. E. Meek, 9 May 1903. PARATYPES: FMNH 4479
(25), CAS-SU 22341 (3), CAS 33825 (old IU 10914) (4), paratopotypes; FMNH 4495 (3).
REMARKS.-Although the jar contains no indication that the four specimens catalogued as IU
10914 are paratypes, they are here so regarded as locality and collector (Meek) are both correct.
Counts for holotype: teeth 4-4; anal rays 9. Counts for CAS-SU paratypes: anal rays 9 (1), 10 (2).
Carl Hubbs (1954:293) downgraded this form from a full species to a subspecies of N. lutrensis,
a decision followed more recently by Contreras (1975). Original description date (31) August
1904.
= Notropis lutrensis forlonensis (Meek)
BULLETIN FLORIDA STATE MUSEUM
Vol. 23, No. 1
Moniana formosa Girard 1857
Girard 1857:201. TYPE LOCALITY: Mimbres R., near Deming, Luna Co., New Mexico. (See
REMARKS below). SYNTYPES: USNM 114 (one set of pharyngeal arches in bone collection), ANSP
3056 (ex USNM 114) (1, 40.0), MCZ 1686 (ex USNM 114) (2, 35.7), John H. Clark, 1851. Girard
(1859a:58) also listed USNM 113 as an original type number.
REMARKS.-The type locality for this species has generally been regarded as the Rio Mimbres
(basin of Laguna de Guzman), in Chihuahua, Mexico (Girard 1859a:58), hut T. Uyeno and R. R.
Miller (unpub. ms), after studying Emory's (1857:46) account and consultation with Dr. W. J.
Koster (Univ. of New Mexico), agreed with him that the type locality is in New Mexico, as indi-
cated above. They also considered the collector and date of collection to be different from that
indicated by Girard (1859a:58) (Dr. C. B. Kennerly, 1854). Girard (1859a:58; pl. 30, figs. 5-8)
listed 24 original syntypes (USNM 113 [4] and USNM 114 [20]) and illustrated one of these.
Counts for USNM 114: teeth 4-4. Counts for ANSP 3056: teeth 44; anal rays 8. Counts for MCZ
1686: teeth 4-4 in both; anal rays 9 in both. Condition of types good for old specimens. This form
is close to N. lutrensis, but was listed as a valid species by Koster (1957:67-68), Bailey et al.
(1970:22), and Minckley (1973:138-139). Contreras (1975) subsequently reviewed this form and
regarded it as a subspecies of N. lutrensis. Original description date 25 April 1857.
= Notropis lutrensis fornosus (Girard)
Alburnus fonnosus Putnam 1863
Putnam 1863:9. TYPE LOCALITY: Mobile, Alabama. LECTOTYPE: UMMZ 162442 (43.0), Judge
LeSene and Albert Stein. Bailey and Suttkus (1952:15) designated lectotype. LECTOPARATYPES:
UMMZ 87183 (6), MCZ 1845 (24), paratopotypes.Bailey and Suttkus identified one other syn-
type (UMMZ 86863) as N. signipinnis.
REMARKS.-This is a senior synonym of N. hypselopterus. Giinther (1868:255) proposed the
substitute name hypselopterus for this species, as Alburnus formosus Putnam 1863 was preoccu-
pied in the genus Leuciscus by Moniana fonnosa Girard 1857. Original description date 28 April
1863.
= Notropis hypselopterus hypsclopterus (Ginther)
Hybopsis fretensis Cope 1869
Cope 1869:382. TYPE LOA:ALITY: near Detroit, Michigan. HOLOTYPE: ANSP 2840 (44.0), E. D.
Cope.
REMARKs.-The type of this species is apparently unique, and thus is regarded as the holo-
type. Fowler (1910: pl. 15, fig. 3) illustrated "type"; calculated length 44.7 mm SL. Hubbs
(1926:45) was first to synonymize this form with N. spilopterus. Original description date (31)
August 1869.
= Notropis spilopterus (Cope)
Moniana frigida Girard 1857
Girard 1857:200. TYPE LOCALITIES: Salado Cr., just E of San Antonio, Bexar Co., Texas; Rio
Sabinal, at Sabinal, Uvalde Co., Texas; and Rio Medina, at Castroville, Medina Co., Texas (all
tributaries of Rio San Antonio); Rio Nueces, ca. 7 mi. W of Uvalde, Uvalde Co., Texas; and Rio
Frio, ca. 10 mi. NE of Uvalde, Uvalde Co., Texas (tributaries of Rio Nueces). According to Hubhs
and Ortenburger (1929b:74), Jordan and Evermann (1896:271) virtually restricted type locality
to Rio Frio, but none of extant syntypes are from there. Also, USNM 124, which was the only
type number given by Jordan and Evermann (op. cit.), represents the series from the Rio Medina.
The type locality, therefore, should not yet he considered firmly fixed. SYNTYPES: USNM 98 (9,
27.347.0) (Rio Sabinal), USNM 100 (three pharyngeal arches) (Salado Creek), USNM 123 (93,
35.9-59.1), ANSP 5062-5063 (presumably ex USNM 123; label in jar says S.I. 2978) (2, 53.0-57.0),
USNM 124 (47, 27.8-61.3 + one set of pharyngeal arches in hone collection; also four specimens
of N. colucellus and one specimen of Moxostonu sp. [probably congestum]) (Rio Medina), John
GILBERT: NOTROPIS TYPE CATALOGUE
H. Clark, 1851. Specimens from Rio Nueces not found. One syntype from Rio Frio (USNM 99:
Dr. C. B. Kennerly, 1853) not found.
REMARKS.-Girard (1858:276-277: pl. 59, figs. 16-20) listed and illustrated one syntype (USNM
99) from Rio Frio. Girard (1859a:56-57; pl. 30, figs. 17-20) listed a total of 133 syntypes (and il-
lustrated one) from three localities as follows: USNM 98 (10 specimens from Rio Sabinal), USNM
1(X) (3 from Salado Creek), USNM 123 (1(X) from Salado Creek), and USNM 124 (20 from Rio
Medina). Condition of types fairly good for old specimens. Counts for USNM 1(): pharyngeal
teeth 4-4 (3). Count for USNM 124: pharyngeal teeth 4-4 (set of arches in hone collection). Jor-
dan (1885a:125) synonvmized this form with V. leonin us, and Hubbs and Ortenburger (1929b:74)
synonymized both with N. lutrensis. Original description date 25 April 1857.
= Notropis lutrensis (Baird and Girard)
Leuciscus frontalis Agassiz 1850
Agassiz 1850:368. TYPE LOCALITY: Montreal R. (trib. to east side of Lake Superior), N of Sault
Ste. Marie, Ontario, Canada. SYNTYPES: MCZ 1751 (1, 116.0), BMNH 1867.4.12.3 (1, 69.0), L.
Agassiz, 5 July 1848.
REMARKS.- Hubbs (1926:46) regarded this form as a subspecies of V. cornutus. Gilbert (1961a)
reduced it to the synonymy of that species. Condition of MCZ type (a tuberculate male) excellent.
Original description date (31 December) 1850.
= Notropis cornutus (Mitchill)
Notropis fimeus Evermann 1892
Evermann 1892:81. TYPE LO(CALITY: Hunter Cr., ca. 9 mi. W of Houston, Harris Co., Texas.
LECTOTYPE: USNM 45558 (41.8), B. W. Evermann, J. T. Scovell, and R. R. Gurley, 20 Novem-
her 1891. Evermann and Kendall (1894: pl. 18) designated lectotype by illustrating "type." Label
in type jar says "specimen drawn." LECTOPAHATYPES: USNM 125072 (ex USNM 45558) (3), para-
topotypes.
REMARKs.-Snelson (1973) reviewed species. Original description date 25 May 1892.
= Sotropis fuimens Evermann
Hypsilepis galactu rus Cope 1868
Cope 1868a:160. TYPE LOCALITY: Holston River system, Virginia. (See REMARKS). LECTOTYPE:
USNM 14981 (83.8), E. D. Cope, October 1867. Gibbs (1961:339) designated lectotype. LECTO-
PARATYPES: ANSP 3381-3444 (65), MCZ 32255 (1), MNHN 4853 (1), paratopotypes.
REMARKs.-Gibbs (1961) reviewed species. Although Gibbs (1961:339) indicated the type lo-
cality as in North Carolina, this is considered erroneous, as Cope did not collect from that part
of the Holston system. Furthermore, the USNM specimen almost certainly came originally from
the ANSP type series, for which Gibbs (op. cit.) gave the locality "Holston River, Virginia." Origi-
nal description date 11 May 1868.
= Notropis galacturus (Cope)
Notropis garmani Jordan 1885
Jordan 18851:813. TYPE LOCALITY: Lago del Mnerto, Parras, Coahuila, Mexico. SYNTYPES:
MCZ 24891 (2, 53.0-56.0), MCZ 24892 (20, 27.8-54.0) UMMZ 86663 (ex MCZ 24892) (6, 40.0-
56.0), USNM 120257 (6, 35.5-51.3), Dr. E. Palmer, 1880.
REMARKs.-Jordan (18851:813) proposed this as a substitute name for Cyprinella rubripiinn
Garman 1881, which was preoccupied in Notropis by Argyreus rubripinnis Heckel 1843. Accord-
ing to Article 72d, International Code of Zoological Nomenclature, the types for Cyprinella
rubripinna must also serve as the types for the replacement name. Species close to N. Iutrensis.
This form has heretofore been considered to be a full species, but Contreras (1975) now regards
it as a subspecies of N. lutrensis. Types in very good condition. Counts for MCZ 24891: teeth 4-4
in both; anal rays 9 (smaller specimen), 10 (larger specimen). Counts for MCZ 24892: anal rays
1978
BULLETIN FLORIDA STATE MUSEUM
9 (7), 10 (12), 11 (1). Counts for USNM 120257: teeth 4-4 (2); anal rays 9 (1), 10 (3), 11 (2). Origi-
nal description date 2 October 1885.
= Notropis lutrensis gannani Jordan
Notropis gennanus Hay 1887
Hay 1887:252-253. TYPE LOCALITY: Smoky Hill River, Wallace, Wallace Co., Kansas. HOLO-
TYPE: USNM 37949 (48.7), 0. P. Hay and M. J. Thompson, July 1885.
REMARKS.-Hubbs (1951b) determined this form to be based on an intergeneric hybrid. Orig-
inal description date 2 July 1887.
= Hybrid: Hybognathus hankinsoni X Notropis heterolepis
Leuciscus gibbosus Storer 1845
Storer 1845:48. TYPE LOCALITY: Tuscaloosa, Alabama. TYPEs: None located.
REMARKS.-Original description inadequate for positive identification. Gilbert (1964:167)
listed it as a possible senior synonym of N. chrysocephalus isolepis. Original description date
(31 December) 1845.
= Status uncertain. Possibly Notropis chrysocephalus isolepis Hubbs and Brown
Moniana gibbosa Girard 1857
Girard 1857:201. TYPE LOCALITY: trib. to Rio Grande, Brownsville, Cameron Co., Texas.
SYNTYPE: USNM 31159 (1, 47.2), Capt. Van Vliet, 1852.
REMARKS.-Girard (1859a:59, pl. 30, figs. 9-12) listed three original syntypes (USNM 105)
and illustrated one of them. Counts for USNM 31159: teeth 4-4; anal rays 9. Jordan (1885a:126)
was first to synonymize this form with N. lutrensis. Contreras (1975) regarded it as a junior syn-
onym of N. I. lutrensis. Original description date 25 April 1857.
= Notropis lutrensis lutrensis (Baird and Girard)
Notropis gibbsi Howell and Williams 1971
Howell and Williams 1971:55-64, figs. 1 (two photographs), 2 a, e, and g. TYPE LOCALITY:
Enitachope Cr., trib. to Hillabee Cr., 3.2 mi. SSW of Ashland, along st. hwy. 9, Clay Co., Ala-
bama. HOLOTYPE: USNM 204933 (74.0), J. D. Williams and H. Harima, 13 July 1968. PARATYPES:
USNM 204934 (40), UMMZ 189199 (45), CU 64311 (45), UAIC 3020 (30), paratopotypes: TU
61102 (28), UF 15933 (28), FSU 16881 (28).
REMARKS.-Species close to N. trichroistius. Original description date 8 March 1971.
= Notropis gibbsi Howell and Williams
Hypsilepis cornutus gihhbus Cope 1868
Cope 1868a:158. TYPE LOCALITY: "southeastern Michigan." TYPEs: None located.
REMARKs.-Hubbs (1926:46) synonymized this form with N. cornutus chrysocephalus. Gilbert
(1964:160) showed that it could also have been based on a hybrid. Original description date 11
May 1868.
= Notropis chrysocephalus chrysocephalus (Rafinesque), or possibly a hybrid between that
species and Notropis cornutus (Mitchill)
Notropis gilbert Jordan and Meek 1885
Jordan and Meek 1885:4-5. TYPE LOCALITY: Village Cr. (trib. to Des Moines R.), SW of Ot-
tumwa, Wapello Co., Iowa. SYNTYPES: USNM 35840 (1, 38.0) (=N. cornutus), UMMZ 61487
(2, 42.0) (=N. dorsalis), UMMZ 61998 (2, 35.0-47.0) (=N. stramineus), ANSP 4137-4143, 4176
(8, 40.0-51.0) (= N. dorsalis), CAS-SU 1456 (9 total: 6, 43.5-45.3 [=N. dorsalis]; 3, 41.0-46.2
[=N. stramineus]), CAS 13518 (ex IU 4612) (15 total: 14, 34.5-46.1 [= N. dorsalis]; 1. 36.1 [=N.
stramineus]), D. S. Jordan and S. E. Meek, August 1884.
REMARKS.-Fowler (1910: pl. 16, fig. 12) illustrated "cotype" (ANSP 4137). Hubbs and Greene
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
(1928:380) were first to synonymize this form with N. dorsalis. Original description date 6 May
1885.
= Notropis dorsalis (Agassiz)
Notropis girardi Hubbs and Ortenburger 1929
Hubbs and Ortenburger 1929a:32-33. TYPE LOCALITY: Cinmnaron R., 3 mi. N\V of Kenton,
Cimmaron Co., Oklahoma. HOLOTYPE: UMMZ 80439 (51.0), A. I. Ortenburger et al., 2-7 July
1926. PARATYPES: UMMZ 80440 (94 originally, now 87), USNM 117564 (ex UMMZ 80440) (5),
BMNH 1933.1.24.1-2 (ex UMMZ 80440) (2), UOMZ 6211 (9)), paratopotypes; UMMZ 73019
(50), UMMZ 73042 (4), UMMZ 80416 (1), UMMZ 80426 (96 originally, now 91), MCZ 36082
(ex UMMZ 80426) (5), UMMZ 80460 (16), UMMZ 80464 (7), UMMZ 80465 (1181, now 1180),
MCZ 32697 (ex UMMZ 80465) (1), UMMZ 80475 (1), UMMZ 80494 (6), UOMZ 6196 (1), UOMZ
6205 (215 ), UOMZ 6217 (4), UOMZ 6252 (1000 ).
REMARKS.-Original description date 16 March 1929.
= Notropis girardi Hubbs and Ortenburger
Leuciscus gracilis Agassiz 1850
Agassiz 1850:370. TYPE LO CALITY: "Lake Huron." SYNTYPES: MCZ 1752 (2, 102.0-110.0), L.
Agassiz, August 1848.
REMARKS.-Jordan and Gilbert (1883:187) were first to svnonvmize this form with N. cornu-
tus and, as first revisers, selected name frontalis over gracilis for the northern subspecies. Con-
dition of types good, though slightly soft. Original description date (31 December) 1850.
= Notropis cornutus (Mitchill)
Hybopsis gracilis Agassiz 1854
See pages 21, 90-91.
Moniana gracilis Girard 1857
Girard 1857:201. TYPE LOCALITY: Acapulco, near Monterrey, Nuevo Leon, Mexico. SYN-
TYPES: USNM 116 (4, 35.0-36.0), ANSP 2992 (ex USNM 116) (1, 57.5), MCZ 1796 (ex USNM 116)
(1, 32.2), Lt. D. N. Couch, winter 1852-1853.
REMARKS.-Girard (1859a:59) listed seven original syntypes. Fowler (1910: pl. 17, fig. 18) il-
lustrated ANSP "cotype." Condition of USNM types good (tuberculate specimens); condition
of ANSP type poor (soft). Counts for ANSP type: teeth 1,4-4,0; anal rays 9. Counts for MCZ type:
teeth 4-4; anal rays 8 (rays difficult to count). Jordan (1885a:125) was first to synonymize this
form with N. lutrensis. Original description date 25 April 1857.
= Notropis lutrensis lutrensis (Baird and Girard)
Photogenis grandipinnis Jordan 1877
Jordan 1877e:62-63. TYPE LOCALITY: Flint R., Georgia. SYNTYPES: None located. The type
number listed in the original description is USNM 9296 ("numerous small specimens in poor
condition," coll. Hugh M. Neisler), but the types of Semotilus thoreauanus are now catalogued
under that number.
REMARKS.-Jordan and Gilbert (1883:185) were first to synonymize this form with N. hyp-
selopterus. Original description date (31 December) 1877.
= Notropis hypselopterus hypselopterus (Giinther)
Notropis greenei Hubbs and Ortenburger 1929
Hubbs and Ortenburger 1929b:78-81. TYPE LOCALITY: Elk R. (trib. to Neosho R.), 7 mi. N of
Grove, Delaware Co., Oklahoma. HOLOTYPE: UMMZ 80978 (42.0), A. I. Ortenburger et al., 11
July 1927. PARATYPES: UMMZ 80979 (4), UOMZ 7733 (2), paratopotypes; UMMZ 60288 (1),
UMMZ 80980 (2), UOMZ 7501 (2), FMNH 1511 (37).
REMARKs.-Hubbs and Ortenburger (1929b:78) listed only four paratopotypes, although a total
BULLETIN FLORIDA STATE MUSEUM
Vol. 23, No. 1
of six are now present in the UMMZ and UOMZ collections. Original description date 16 March
1929.
= Notropis greenei Hubbs and Ortenburger
Cyprinella gunnisoni Girard 1857
Girard 1857:197. TYPE LOCALITY: Cottonwood R., ca. 5 mi. NW of Durham, Marion Co.,
Kansas. (See REMARKS). SYNTYPES: USNM 139 (24, 25.4-54.0 + one set of pharyngeal arches
in bone collection), F. Creutzfeldt, 3-4 July 1853.
REMARKs.-Girard (1858:267) listed only 12 original syntypes. Counts for types: teeth 4-4 (1),
0,4-4,1 (1, arches in bone collection); anal rays 9 (5). Types in very poor condition (desiccated).
Jordan (1885a:123) synonymized this form with N. hubalinus and Hubbs and Ortenburger
(1929a:34; 1929b:75) synonymized both with N. lutrensis.
The type localities for this and for two other species (Cyprinella ludibunda and C. lugubris),
apparently collected simultaneously, have been confused. Girard (1857:197; 1858:271-272) gave
the type locality as "Cottonwood Creek, Utah." Jordan and Evermann (1896:273) determined
this to be in error, but gave no alternative possibility. Snyder (1921:27) suggested that this prob-
ably was a tributary of the Arkansas River, in Kansas; but Jordan, Evermann, and Clark (1930:130)
"corrected" this to Cottonwood Creek, tributary to the Rio Grande, in Colorado. Based on Beck-
with's (1855:16) account, table of distances (op cit: 115), the map (no. 2) appearing in volume 11
of the Railroad Survey reports, and the fish species collected, I believe (as did Snyder [op. cit.])
that this is the Kansas locality listed above.
Original description date 25 April 1857.
= Notropis lutrensis (Baird and Girard)
Cyprinus haematopterus Rafinesque 1820
Rafinesque 1820a:6. TYPE LOCALITY: Hudson River system, New York. TYPES: None located.
REMARKs.-Gilbert (1964:141) synonymized this form with N. cornutus. Original description
date 1 March 1820.
= Notropis cornutus (Mitchill)
Notropis harper Fowler 1941
Fowler 1941:231-233, figs. 5-6. TYPE LOCALITY: small spring, ca. 220 yds. SE of Manatee
Spring, off Suwannee R., ca 7 mi W of Chiefland, Levy Co., Florida. HOLOTYPE: ANSP 69151
(32.0), Francis Harper, 24 June 1939. PARATYPES: ANSP 69152 (1) paratopotype; ANSP 69153-
69157 (5).
REMARKS.-This species usually, but not always, has a barbel at each corner of the month.
Herald and Strickland (1949:102) therefore referred it to the genus Erimystax, presumably on the
advice of C. L. Hubbs. Bailey (1951:192) downgraded Erimystax to a subgenus of Hybopsis, in
which species harper remained until returned to Notropis by Gilbert and Bailey (1972:11). Hubbs
and Crowe (1956:5) recognized two subspecies of Hybopsis harper subterraneana and the nomi-
nate form), but unpublished data (Howell 1960) indicate these should not be so considered. Orig-
inal description date 18 March 1941.
= Notropis harper Fowler
Notropis henryi Fowler 1942
See page 22.
Alburnops heterodon Cope 1865
Cope 1865a:281. TYPE LOCALITY: Two localities listed in original description: Lansing and
Grosse Isle, Michigan. TYPES: None located. Types lost, according to Fowler (1918:20).
REMARKS.-Original description date 13 February 1865.
= Notropis heterodon (Cope)
GILBERT: NOTROPIS TYPE CATALOGUE
Notropis heterolepis Eigenmann and Eigenmann 1893
Eigenmann and Eigenmann 1893a:152. TYPE LOCALITY: trib. to Qu'Appelle R. at Fort Qu'-
Appelle (50046'N, 103048'W), ca 2 mi SW of Melville, Saskatchewan, Canada. HOLOTYPE:
BMNH 1892.12.30.532 (47.7), C. H. Eigenmann, August or September 1892.
REMARKS.-This species was long erroneously called N. cayuga (Jordan and Evermann
1896:260 [In part]; Forbes and Richardson 1920:133, pl. opp. p. 128, lowest fig.). Hubbs (1926:41)
was first to clarify its status. Original description date 4 February 1893.
= Notropis heterolepis Eigenmann and Eigenmann
Notropis horatii Cockerell 1911 emendedd spelling Notropis horacei)
Cockerell 1911:614-615. TYPE LOCALITY: small trib. (presumably) to S Platte R., near Jules-
burg, Sedgwick Co., Colorado. HOLOTYPE: USNM 70313 (43.5), Horace G. Smith.
REMARKS.--N. horacei proves to be a junior synonym of N. dorsalis, despite Cockerell's con-
tention to the contrary. Counts for holotype: teeth 1,4-4,1; dorsal and anal rays both 8; lateral-
line scales ca. 36; body circumference scales ca. 14-2-13; caudal-peduncle circumference scales
5-2-5. The type locality is in an area of intergradation between the presumed subspecies N. d.
dorsalis and N. d. piptolepis. The species was named for its collector, Horace G. Smith. Original
description date 3 November 1911.
= Notropis dorsalis (Agassiz)
Clupea hudsonia Clinton 1824
Clinton 1824:49-50, fig. 1. TYPE LOCALITY: Hudson R., New York. TYPES: None located.
REMARKs.-Seaman (1968) reviewed species. This widely distributed, variable species com-
prises two distinct morphological types, both of which occur in the Hudson River system, New
York. As this is the type locality of Clupea hudsonia, designation of a neotype seems desirable.
Original description date (31 December) 1824.
= Notropis hudsonius (Clinton)
Ceratichthys hyalinus Cope 1868
See pages 21, 91.
Leuciscus hypselopterus Cinther 1868
Giinther 1868:255. TYPE LOCALITY: Mobile, Alabama. LECTOTYPE: UMMZ 162442 (43.0),
Judge LeSene and Albert Stein. Bailey and Suttkus (1952:15) designated lectotype. LECTOPARA-
TYPES: UMMZ 87183 (6), MCZ 1845 (24), paratopotypes. Bailey and Suttkus identified one other
syntype (UMMZ 86863) as N. signipinnis.
REMARKs.-Giinther (1868:255) proposed this as a substitute name for Alburnus formosus Put-
nam 1863, which was preoccupied in the genus Leuciscus by Moniana formosa Girard 1857.
The types of L. hypselopterus are thus the same as for A. formosus. Original description date 14
March 1868.
= Notropis hypselopterus hypsehlpterus (Giinther)
Notropis hypsilepis Suttkus and Raney 1955
Suttkus and Raney 1955b:161-170, figs. 1-2. TYPE LOCALITY: trib. to Soque R., 0.4 mi W of
Soque R. and 1.5 mi W of Clarksville, on st. hwy. 115, Habersham Co., Georgia. HOLOTYPE: CU
17442 (51.0), E. C. Raney, R. D. Suttkus, R. H. Backus, and C. R. Robins, 1 April 1950. PARATYPES:
CU 28237 (6), paratopotypes; CU 17138 (19), CU 17279 (3), CU 18102 (8), CU 19625 (9), CU
21138 (1), UMMZ 136086 (42 originally, now 39), UF 14610 (ex UMMZ 136086) (3), UMMZ
142910 (2), UMMZ 157957 (1).
REMARKS.-Swift (1970) reviewed species. Original description date 25 February 1955.
= Notropis hypsilepis Suttkus and Raney
Ceratichthys hypsinotus Cope 1870
See pages 21, 91.
BULLETIN FLORIDA STATE MUSEUM
Notropis spilopterus hypsisomatus Gibbs 1958
Gibbs 1958:195-198, fig. 4b-c. TYPE LOCALITY: bay at N end of Wonder Lake, McHenry
Co., Illinois. HOLOTYPE: CU 28541 (68.6), J. N. Layne and Lois V. Linderoth, 2 July 1950. PARA-
TYPES: CU 22305 (2), paratopotypes; CU 17931 (48), CU 17946 (15), CU 17960 (31), CU 17984
(6), CU 22326 (12), CU 22330 (33).
REMARKS.-This is a weakly defined form, differing from the nominate subspecies by slight
differences in lateral-line scale count, body depth, and nuptial-tubercle development on the
lower part of the caudal peduncle. It has been recognized in some publications (Larimore and
Smith 1963:323). Original description date 10 January 1958.
= Notropis spilopterus (Cope)
Alburnops illecebrosus Girard 1857
Girard 1857:194. TYPE LOCALITY: Arkansas R., Fort Smith, Arkansas. LECTOTYPE: ANSP
3034 (58.5), Dr. George C. Shumard, 1853. Lectotype designated by Meek (in Jordan 1885a:123).
LECTOPARATYPES: USNM 66 (18), MCZ 1783 (2), UMMZ 56286 (ex MCZ 1783) (1), FMNH 3142
(1), paratopotypes. In addition, five of the original USNM syntypes represent N. hoops (see Hubbs
and Ortenburger 1929a:29).
REMARKs.-Girard (1858:262-263; pl. 57, figs. 5-8) listed only 24 original syntypes and illus-
trated one of these. Some older references to N. illecebrosus refer to N. boops (Jordan and Ever-
mann 1896:268-269; Forbes and Richardson 1920:140-141, fig. 35), as pointed out by Hubbs
(1926:43) and Hubbs and Ortenburger (1929a:29), who first clarified the status of this species.
Jordan and Gilbert (1883:192-193) synonymized N. illecebrosus and N. shumardi and, as first re-
visers, selected the latter name; however, this was generally overlooked until pointed out by Gil-
bert and Bailey (1962:809). It now appears that Jordan and Gilbert's action, though correct, was
based on the assumption that both names apply to the species now called N. hoops (R. D. Sutt-
kus, in litt). Original description date 25 April 1857.
= Notropis shumardi (Girard)
Notropis imeldae Cort6s 1968
CortBs 1968:185-192, 1 fig. TYPE LOCALITY: Rio Atoyac, Juchatengo (16020'N, 97005'W),
Oaxaca, Mexico. HOLOTYPE: IPN P-915 (59.9), Imelda Martinez and Lauro Gonzalez, 16 April
1965. PARATYPES: UMMZ 188317 (5), TU 55375 (5), 183 other specimens presumably at same
institution as holotype; all paratopotypes.
REMARKS.-This species usually has a small barbel at each corner of the mouth. Original de-
scription date 30 July 1968.
= Notropis imeldae Cortes
Notropis ionthas Jordan and Gilbert 1886
See page 22.
Notropis ipni Alvarez and Navarro 1953
See page 22.
Hypsilepis iris Cope 1875
Cope, in Cope and Yarrow 1875:653-654; pl. 31, figs. 4-4a. Type locality: Rio Grande, at
San Ildefonso, ca 10 mi E of Los Alamos, Santa Fe Co., New Mexico. SYNTYPES: USNM 16976
(116 total originally, now 111 total: 102, 28.2-52.3 [= N. lutrensis]; 9 = Gila pandora), MCZ 35927
(ex USNM 16976) (5, 28.3-31.0) [=N. lutrensis]; USNM 16977 (268 total: 188, 25.0-42.0 [=N.
lutrensis]; I = N. jemezanus; 19 Gila pandora; 50 = Rhinichthys cataractae; 10 = Catostomnus
plebeius), USNM 16980 (98 total: 34, 20.0-31.0 [= N. lutrensis]; 29= Gila pandora; 6= Rhinich-
thys cataractae; 29 = Catostomus plebeius), H. C. Yarrow and E. D. Cope, August 1874.
REMARKS.-Jordan (1885b:812) was first to synonymize this form with N. lutrensis. Contreras
(1975) considered it to be a junior synonym of N. I. lutrensis. Condition of types poor (soft). Counts
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
for MCZ types: anal rays 9 in all. One of the largest specimens in USNM 16977 removed for pos-
sible designation as lectotype; counts for specimen are teeth 1,4-4,1 and anal rays 9. Original
description date (31 December) 1875.
= Notropis lutrensis lutrensis (Baird and Girard)
Notropis cornutus isolepis Hubbs and Brown 1927
Hubbs and Brown, in Ortenburger and Hubbs 1927:129-131. TYPE LOCALITY: Mountain
Fork R., 10 mi SE of Broken Bow, McCurtain Co., Oklahoma. HOLOTYPE: UMMZ 73090 (101.0),
A. I. Ortenburger et al., June-July 1925. PARATYPEs: UMMZ 73091 (4), UOMZ 5971 (5), para-
topotypes; UMMZ 73048 (2), UMMZ 73097 (16 originally, now 14), MCZ 32696 (ex UMMZ
73097) (1), USNM 117561 (ex UMMZ 73097) (1), UOMZ 5969 (1), UOMZ 5970 (38), UOMZ 5972
(1), UOMZ 5973 (2).
REMARKs.-Gilbert (1964:167-172) reviewed this form. It was regarded as a subspecies of N.
cornutus until 1961, when Gilbert (1961a) changed it to a subspecies of N. chrysocephalus.
Menzel (1970, 1976) recommended elevation to a full species, based on analysis of serum protein
characteristics. Wall (1968) and Clark (1974), in their analyses of morphometric and meristic
characters in populations from southern and western tributaries of the lower Tennessee River,
found what appears to be even and consistent intergradation between chrysocephalus and iso-
lepis. This suggests a subspecific relationship, although this problem cannot be finally resolved
until the serum protein patterns of these morphologically intermediate populations are studied.
Original description date 1 January 1927.
= Notropis chrysocephalus isolepis Hubbs and Brown
Alburnellus jaculus Cope 1868
Cope, in Gunther 1868:255-256. TYPE LOCALITY: Walker Cr., either Bland or Giles Co., Vir-
ginia; St. Joseph R. and Dowagiac Cr. (trib. to St. Joseph R.), Berrien Co., Michigan. SYNTYPES:
BMNH 1868.1.10.24 (3, 43.0-58.0) (Walker Cr.), E. D. Cope, August-September 1867; ANSP
2577 (1, 42.0), MCZ 4688 (2, 50.6-55.2) (St. Joseph R. and Dowagiac Cr.), E. D. Cope.
REMARKS.-Although the Michigan localities were not specifically mentioned in the original
description, these are still regarded as type localities and the specimens taken there as syntypes,
for reasons discussed earlier in this paper. The original description specified the range of the
species as including Michigan and Virginia. Jordan and Copeland (1877:154) synonymized this
form with Minnilus dinemus, and Jordan and Evermann (1896:293) synonymized both with N.
atherinoides. I subsequently identified the ANSP and MCZ syntypes of A. jaculus as N. rubellus
(Agassiz). James Chambers (in litt.) confirmed that the syntypes in the BMNH are also this spe-
cies, based on a comparison with fresh specimens of N. rubellus supplied by me. For further dis-
cussion of nomenclature of A. jaculus see account of Leuciscus copii. Original description date
14 March 1868.
= .otropis rubellus (Agassiz)
Episenma jejuna Forbes 1878
Forbes, in Jordan 1878a:60-61. TYPE LOCALITY: Illinois R., at Henry, Marshall Co., Illinois.
The Illinois R., at Pekin, Tazewell Co., Illinois, perhaps should also be considered a type locality
(see below). SYNTYPES: INHS 26947(17, 31.8-58.6), CAS-SU 1278(3, 54.0-60.2) (label in jar says
"Illinois River, Illinois"), CAS 29302 (old IU 4725) (2, 51.8-54.0) (label in jar says simply "Illi-
nois"), Stephen A. Forbes, summer 1877. (Label in jar of INHS types says 1 November 1877).
Also INHS 26946 (2, 55.5-58.9) from the Illinois R., Pekin, Tazewell Co., Illinois, are catalogued
as "types," but label in jar indicates the specimens were collected 2 June 1880, which is subse-
quent to the original description.
REMARKS.-Hubbs (1926:43) was first to synonymize this form with N. blennius. Hubbs and
Bonham (1951:104) and Hubbs and Lagler (1958:82) regarded it as a valid subspecies, but this is
not substantiated by the preliminary findings of Suttkus and Clemmer (1968). Counts for CAS
BULLETIN FLORIDA STATE MUSEUM
29302: teeth 2,4-4,2 and anal rays 7 in both; body circumference scales 13-2-11; caudal-peduncle
circumference scales 5-2-5. Original description date (30) June 1878.
= Notropis blennius (Girard)
Alburnellus jemezanus Cope 1875
Cope, in Cope and Yarrow 1875:650-651; pl. 31, figs. 3-3a. TYPE LOCALITY: Rio Grande, at
San Ildefonso, ca 10 mi E of Los Alamos, Santa Fe Co., New Mexico. SYNTYPEs: USNM 16981
(7, 33.6-57.0), H. C. Yarrow and E. D. Cope, August 1874.
REMARKS.-Jordan (1885b:815) synonymized this form with N. dilectus (=N. atherinoides),
which ,Was followed until Hubbs (1940:6) revalidated the species, a decision based on unpublished
studies by Hubbs and Myron Gordon. Types in rather poor condition. Counts: anal rays 9 (1)
(rays difficult to count), 10 (7); one type with three rows of teeth. Original description date (31
December) 1875.
= Notropis jemezanus (Cope)
Notropis jordanii Eigenmann and Eigenmann 1893 emendedd spelling Notropis jordani)
Eigenmann and Eigenmann 1893b:592. TYPE LOCALITY: S. Saskatchewan R., Medicine Hat,
Alberta, Canada. HOLOTYPE: BMNH 1892.12.30.531(53.5), C. H. Eigenmann, August or Septem-
ber 1892.
REMARKs.-The name jordanii is a substitute name for N. albeolus Eigenmann and Eigen-
mann 1893, preoccupied in Notropis by N. albeolus Jordan 1889. The holotype for this form is
thus the same as for the first N. albeolus above. For further remarks see account of N. albeolus
Eigenmann and Eigenmann. Original description date 15 June 1893.
= Notropis blennius (Girard)
Moniana jugalis Cope 1871
Cope 1871:439-440. TYPE LOCALITY: Missouri R., at St. Joseph, Missouri. (See under RE-
MARKS). LECTOTYPE: ANSP 3144 (51.7) Dr. William Stimpson. Fowler (1910: pl. 17, fig. 20) desig-
nated lectotype by illustrating "type" (ANSP 3144); calculated length 52.5 mm SL. LECTOPARA-
TYPES: ANSP 3145-3150 (16), paratopotypes.
REMARKS.-Jordan (1885b:812) was first to synonymize this form with N. lutrensis. Fowler
(1910: pl. 17, fig. 20) illustrated "type" (ANSP 3144); calculated length 52.5 mm SL. Types in
good condition for old specimens. Although Fowler (1910) listed only 7 syntypes (this number
is also indicated on the jar label), there are actually 17 specimens in the jar. Inasmuch as all are
in the same state of preservation and there is no record to indicate subsequent addition of speci-
mens, one may assume that the original total (7) is in error. Counts for lectotype: teeth 1,4-4,1;
anal rays 9. Type specimens may not have been collected exactly at St. Joseph, Missouri, but
may have come from near there (see REMARKS under Alburnellus percobromus). Original descrip-
tion date (31 December) 1871.
= Notropis lutrensis (Baird and Girard)
Notropis kanawha Jordan and Jenkins 1889
Jordan and Jenkins, in Jordan 1889a:354-355; pl. 44, fig. 5. TYPE LOCALITY: Reed Cr., near
Wytheville, Wythe Co., Virginia. LECTOTYPE: USNM 39928 (58.0), D. S. Jordan, O. P. Jenkins,
B. W. Evermann, and S. E. Meek, summer 1888. Bailey and Gilbert (1960) designated lectotype.
LECTOPARATYPES: USNM 179894 (ex USNM 39928) (3). Three other original syntypes (CAS-SU
624) were reidentified by Bailey and Gilbert (1960:354-355) as N. colucellus.
REMARKS.-Bailey and Gilbert (1960) reviewed this form and determined that it was based on
an interspecific hybrid. Counts for lectotype: teeth 1,44,0; anal rays 9. Original description date
5 July 1889.
= Hybrid: Notropis rubellus X Notropis colucellus
Notropis keimi Fowler 1909
Fowler 1909:533-535, pl. 27. TYPE LOCALITY: trib. of Allegheny R., S of Port Allegany,
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
McKean Co., Pennsylvania. HOLOTYPE: ANSP 31126 (48.0), T. D. Keim and H. W. Fowler, August
1904. PARATYPES: ANSP 24045-24047 (3).
REMARKS.-Huhbs and Greene (1928:381) regarded this form as a synonym of N. dorsalis,
but later Hubbs and Lagler (1947:67; 1958:84) accorded it subspecific status. The former de-
cision is usually followed. Original description date 28 January 1909.
= Notropis dorsalis (Agassiz)
Notropis kendalli Evermann and Cockerell 1909
Evermann and Cockerell 1909:186-187. TYPE LOCALITY: Cross Lake Thoroughfare (stream
connecting Salmon and Cross lakes), Aroostook Co., Maine. HOLOTYPE: USNM 64150 (44.0), Dr.
William C. Kendall, 9 July 1903.
REMARKs.-Hubbs (1926:42) originally synonymized this form with N. bifrenatus, but later
Hubbs and Brown (1929:30) synonymized it with N. heterolepis. Counts for holotype: teeth 4-4;
anal rays 8; predorsal scales 18; caudal-peduncle circumferential scales 7-2-5. Original descrip-
tion date 8 December 1909.
= Notropis heterolepis Eigenmann and Eigenmann
Ceratichthys labrosus Cope 1870
See pages 21, 91.
Hybopsis lacertosus Cope 1868
Cope 1868b:232. TYPE LOCALITY: Bear Cr., trib. to Middle Fk. of Holston R., Virginia. LEC-
TOTYPE: ANSP 2835 (85.0), E. D. Cope. Fowler (1910: pl. 19, fig. 33) designated lectotype by il-
lustrating "type" (ANSP 2835); calculated length 91.2 mm SL. LECTOPARATYPES: ANSP 2836-
2839 (4), paratopotypes.
REMARKS.-Types in extremely poor condition, the specimens partly decomposed. Gilbert
(1964:159-161) synonymized this form with N. chrysocephalus chrysocephalus. Original descrip-
tion date 2 December 1868.
= Notropis chrysocephalus chrysocephalus (Rafinesque)
Moniana laetabilis Girard 1857
Girard 1857:200. TYPE LOCALITY: Hurrah Cr. (trib. to Pecos R.), ca 10 mi NE of Santa Rosa
(35005'23.7"N, 10443'27.3"W), Guadalupe Co., New Mexico. SYNTYPES: USNM 120 (one set
of pharyngeal arches in bone collection), MCZ 1691 (1, 34.5), ANSP 3029 (1, 35.0), H. B. M611-
hausen, 24 September 1853.
REMARKS.-Girard (1858:275) listed 12 original syntypes (USNM 120). Condition of types fair
to poor (discolored but firm). Counts: USNM 120 (teeth 4-4); ANSP 3029 (teeth 4-, anal rays 9);
MCZ 1691 (teeth 44, anal rays 9). Jordan (1885a:125) was first to synonymize this form with
N. lutrensis. Contreras (1975) regarded it as a junior synonym of N. 1. lutrensis. Original descrip-
tion date 25 April 1857.
= Notropis lutrensis lutrensis (Baird and Girard)
Notropis leedsi Fowler 1942
Fowler 1942:2-4, figs. 3-4. TYPE LOCALITY: Ohoopee R., ca 2% mi N of Oak Park, Emanuel
Co., Georgia. HOLOTYPE: ANSP 69969 (67.0), Francis Harper, 1 May 1933.
REMARKS.-Gibbs (1955) reviewed species. Original description date 16 September 1942.
= Notropis leedsi Fowler
Moniana leonina Girard 1857
Girard 1857:199. TYPE LOCALITY: Leon R. (= Leon Cr.), trib. to San Antonio R., Bexar Co.,
Texas. SYNTYPES: USNM 115 (one set of pharyngeal arches in osteological collection), MCZ 1687
(ex USNM 115) (1, 59.8), Dr. C. B. Kennerly, 1853.
REMARKs.-Girard (1858:273-274; pl. 59, figs. 6-10) listed 18 original syntypes (USNM 115)
and illustrated one of these. Condition of MCZ type (a tuberculate male) good for old specimen.
Counts: USNM 115 (teeth 4-4); MCZ 1687 (teeth 44, anal rays 9). Jordan and Evermann
1978
BULLETIN FLORIDA STATE MUSEUM
(1896:272) were first to synonymize this form with N. lutrensis. Original description date 25 April
1857.
= Notropis lutrensis (Baird and Girard)
Cyprinella lepida Girard 1857
Girard 1857:197-198. TYPE LOCALITY: Rio Frio (trib. to Rio Nueces), either Real or Uvalde
Co., Texas. SYNTYPE: USNM 126 (one set of pharyngeal arches in osteological collection), Dr. C.
B. Kennerly, 1853.
REMARKS.-Girard (1858:268-269; pl. 58, figs. 21-25) listed seven original syntypes (USNM
126) and illustrated one of these. Jordan and Evermann (1896:273) synonymized this form with
N. bubalinus, and Hubbs and Ortenburger (1929a:34) synonymized the latter with N. lutrensis.
Clark Hubbs (1954b:281-283, fig. 2) recognized N. lepidus as a valid species, and it was so listed
by Bailey et al. (1970:22). Later he (Clark Hubbs 1972) relegated it, without comment, to the
synonymy of N. lutrensis, but Lytle (1972) regarded it as a valid species most closely related to
N. proserpinus. Counts for type: teeth 4-4. Original description date 25 April 1857.
= Notropis lepidus (Girard)
Alburnus lepidulus Girard 1857
Girard 1857:212. TYPE LOCALITY: Black Warrior R., Alabama. SYNTYPES: None located.
Specimens collected by Prof. A. Winchell.
REMARKS.-Jordan and Evermann (1896:294) synonymized this form with N. dilectus, and
Hubbs and Ortenburger (1929b:83) later synonymized dilectus with N. atherinoides. Smith-Vaniz
(1968:43) also regarded lepidulus as a probable synonym of N. atherinoides. Original description
date 25 April 1857.
= probably Notropis atherinoides Rafinesque
Notropis lermae Evermann and Goldsborough 1902
Evermann and Goldsborough 1902:147-148, fig. 3. TYPE LOCALITY: Lake Lerma, near town
of Lerma, state of Mexico, Mexico. HOLOTYPE: USNM 50003 (47.7), Dr. J. N. Rose, 9 August
1901. PARATYPES: USNM 50004 (32), USNM 126648 (formerly USFC reserve 977) (10), CAS-SU
6762 (10), paratopotypes. The last series was not listed in Bohlke's (1953) type catalogue.
REMARKS.-Counts for holotype: teeth 4-; anal rays 7; lateral-line scales 41. Original descrip-
tion date 3 May 1902.
= Notropis sallei (Giinther)
Photogenis leuciodus Cope 1868
Cope 1868a:165. TYPE LOCALITY: tribs. of Holston River, Virginia. LECTOTYPE: ANSP 2336
(62.0), E. D. Cope, Fowler (1910: pl. 20, fig. 45) designated lectotype by illustrating "type
(ANSP 2336); calculated length 62.4 mm SL. LECTOPARATYPES: ANSP 2337-2358 (22), MCZ
25161 (1). One specimen (ANSP 2359), representing either N. chrysocephalus or N. coccogenis,
removed from original ANSP syntypic series and recatalogued.
REMARKS.-Types in fair to poor condition. Counts for lectotype: teeth 1,4-4,1; anal rays 8.
Counts for ANSP lectoparatypes: teeth 1,4-4,1 (16); anal rays 8 (20), 9 (1). Counts for MCZ lec-
toparatype: teeth 1,4-4,1; anal rays 8. Original description date 11 May 1868.
= Notropis leuciodus (Cope)
Photogenis leucops Cope 1868
Cope, in Gunther 1868:252. TYPE LOCALITY: Sinking Cr. tribe to Kanawha [= New] R.), "near
Austinville, Wythe Co.," Virginia. (See REMARKS below). LECTOTYPE: ANSP 2581 (83.0), E. D.
Cope, August-September 1867. Fowler (1910: pl. 21, fig. 52) designated lectotype by illustrating
"type" (ANSP 2581); calculated length 85.6 mm SL. This antedates Gilbert's (1971a:478) sub-
sequent lectotype designation, which presumably was based on the same specimen. LECTOPARA-
TYPES: ANSP 2582-2628 (46), ANSP 3017-3024 (8), ANSP 3063-3080 (20), BMNH 1868.1.10.10
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
(4), MNHN 4854 (1), paratopotypes. Two other original syntypes (ANSP 2629-2630) have been
recatalogued as N. spilopterus and N. rubellus, respectively.
REMARKS.--Photogenis leucops was originally described, as a junior synonym of Leuciscus
photogenis, in Gunther (1868:252), but the name is available under the provision of Article 11D
of the International Code of Zoological Nomenclature (see Gilbert 1971a:476). Jordan (1878d:
297-298) was first to synonymize this form with N. photogenis. Gilbert (1971a:477-478) discussed
the nomenclature of this form and status of the types. The location of Sinking Creek was given in
the original description as "near Austinville, Wythe Co." Unless this stream is different from the
one known by that name today, the type locality is neither close to Austinville nor is it in Wythe
County (R. E. Jenkins, pers. comm.). Original description date 14 March 1868.
= Notropis photogenis (Cope)
Photogenis leucopus Jordan and Brayton 1878
Jordan and Brayton 1878:41-42. TYPE LOCALITY: Chattahoochee R., at Shallow Ford, NW
of Gainesville, Hall Co., Georgia. LECTOTYPE: USNM 31124 (77.8), D. S. Jordan and C. H. Gilbert,
summer 1877. Lectotype designated by Bailey and Gibbs (1956:2). LECTOPARATYPES: USNM
163963 (1), MCZ 35935 (1), paratopotypes.
REMARKs.-Jordan and Evermann (1896:277) synonymized this form with N. eurystomus;
Bailey, Winn, and Smith (1954:128) synonymized eurystomus with N. cenustus; and Gibbs
(1957:189-190) synonymized both with N. cenustus cercostigma. Original description date (31
December) 1878.
= Notropis cenustus cercostigma (Cope)
Hybopsis lineapunctata Clemmer and Suttkus 1971
See pages 21, 91-92.
Alburnus lineolatus Agassiz 1863
Agassiz, in Putnam 1863:9. TYPE LOCALITY: Osage R., Missouri. SYNTYPES: None located.
Specimens collected by George Stolley in 1854 supposedly at MCZ. A specimen (44.0 mm SL)
in the British Museum (BMNH 1867.4.12.15), which was sent by the MCZ to the Liverpool Mu-
seum and later transferred to the BMNH, likely is a syntype, but this cannot be established posi-
tively.
REMARKS.-C. L. Hubbs identified the above BMNH specimen as N. delicious auct. (=N.
stramineus) in 1958. Suttkus (1958:313) discussed status of this specimen, and listed Alburnus
lineolatus as a possible senior synonym of N. stramineus. Original description date 28 April 1863.
= a probable senior synonym of Notropis stroamineus (Cope)
Nototropis lirus Jordan 1877
Jordan 1877c:342. TYPE LOCALITY: tribs. of Etowah R., near Rome, Floyd Co., Georgia. LEC-
TOTYPE: ANSP 19842 (41.5), D. S. Jordan and C. H. Gilbert, July 1876. Lectotype herein desig-
nated by F. F. Snelson, Jr. LECTOPARATYPES: USNM 17876 (6), USNM 20138 (2), USNM 101158
(5), BMNH 1880.1.21.7-8;24 (3), MNHN A. 1303 (3).
REMARKs.-Snelson (1970) reviewed species. Name first appeared as a nomen nudum (Lyth-
rurus lirus) in Jordan and Copeland (1877:152). Fowler (1910: pl. 21, fig. 55) illustrated "cotype."
Original description date (30) June 1877.
= Notropis lirus (Jordan)
Hybopsis longiceps Cope 1868
Cope 1868b:231. TYPE LOCALITIES: headwaters of Roanoke R., Montgomery Co., Virginia;
and headwaters of James R., Giles Co., Virginia. LECTOTYPE: ANSP 4108 (58.0), E. D. Cope,
July 1867. Fowler (1910: pl. 16, fig. 9) designated lectotype by illustrating "type" (ANSP 4108);
calculated length 56.5 mm SL. LECTOPARATYPES: ANSP 4109-4122 (14). Three additional original
syntypes reidentified as N. colucellus and recatalogued under that name.
REMARKS.-This form was erroneously synonymized by Jordan and Copeland (1877:150) with
BULLETIN FLORIDA STATE MUSEUM
Hybopsis (=Notropis) microstomus, and by Jordan (1885b:811) with N. delicious (=N. stra-
mineus). Jordan and Evermann (1896:264) were first to synonymize it with N. procne. Raney
(1947) considered it to be a valid subspecies of N. procne, but Snelson (1971:461) indicated this
may not be so. Original description date 2 December 1868.
= Notropis procne (Cope)
Alburnops longirostris Hay 1881
Hay 1881:504. TYPE LOCALITY: Chickasawha R., Enterprise, Clarke Co., Mississippi. SYN-
TYPES: USNM 27440 (11, 37.0-43.2), USNM 101118 (24, 30.0-47.2), O. P. Hay, March-April 1880.
REMARKS.-Original description date 23 February 1881.
= Notropis longirostris (Hay)
Notropis louisae Fowler 1940
Fowler 1940:2-4, color pl. 1 (left fig.). TYPE LOCALITY: stream near Punta Gorda, Charlotte
Co., Florida. HOLOTYPE: ANSP 69131 (43.5), C. C. G. and L. C. Chaplin, May 1940. PARATYPES:
ANSP 69132-69140 (9).
REMARKs.-Bailey, Winn, and Smith (1954:129) were first to synonymize this form with N.
maculatus. Original description date 3 October 1940.
= Notropis maculatus (Hay)
Notropis louisianae Evermann 1898
Evermann, in Jordan and Evermann 1898:2801. TYPE LOCALITY: Atchafalaya R., Melville,
St. Landry Psh., Louisiana. HOLOTYPE: USNM 48902 (58.0), Fred M. Chamberlain, 5 May 1897.
PARATYPE: USNM 125607 (1), paratopotype.
REMARKS.-This form apparently has not heretofore been formally synonymized with N.
atherinoides. Original description date (30) November 1898.
= Notropis atherinoides Rafinesque
Luxilus lucidus Girard 1857
Girard 1857:203. TYPE LOCALITY: 20 mi. W of Choctaw Agency, Oklahoma. This locality is
either San Bois Cr. or a branch thereof, 20 mi W of Scullyville, Haskell Co., Oklahoma. (See
under REMARKS). LECTOTYPE: ANSP 4536 (ex USNM 56) (50.0), H. B. M6llhausen, 31 July-1
August 1853. Lectotype designated by Snelson and Pflieger (1975:233). LECTOPARATYPES: ANSP
4537 (1) (Snelson and Pflieger [1975:233] erroneously listed catalogue number as ANSP 4547),
MCZ 1782 (1), paratopotypes.
REMARKs.-Girard (1858:282; pl. 60, figs. 9-12) listed a total of 24 original syntypes from two
localities (12 from USNM 55 [Coal Cr., trib. to S. Fk. of Canadian R., Oklahoma] and 12 from
USNM 56 [W of Choctaw Agency]) and illustrated one of these. As the lectotype was originally
part of USNM 56, the type locality is restricted on that basis. Locality data with ANSP 4536-
4537 says "Coal Creek," but as the specimens were said to have come from USNM 56, these data
are assumed to be in error. (For further discussion see Snelson and Pflieger [1975:233-234]). Loca-
tion of type locality and date of collection were determined by reference to Whipple's (1854:10-
12) itinerary of the expedition on which the types were taken and from the localities listed in Ap-
pendix D (Whipple 1856:70-71). Jordan (1885a:122) was first to synonymize this form with N.
umbratilis (Girard 1857), and in so doing selected the name umbratilis; he thus qualifies as first
reviser. Original description date 25 April 1857.
= Notropis umbratilis umhratilis (Girard)
Cyprinella ludibunda Girard 1857
Girard 1857:199. TYPE LOCALITY: Cottonwood R., ca 5 mi NW of Durham, Marion Co.,
Kansas. SYNTYPES: ANSP 2841 (ex USNM 132) (1, 31.5) (= Notropis stramineus), MCZ 1799 (ex
USNM 132) (1, 30.0) (= Notropis volucellus), F. Creutzfeldt, 3-4 July 1853.
REMARKS.-For discussion of correct type locality see REMARKS under Cyprinella gunnisoni.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
Girard (1858:271-272) listed seven original syntypes (USNM 132). Cyprinella Ildlibunda is a
senior synonym of both N. straoiineus and N. volucellus, a fact that was overlooked bv Suttkus
(1958). If one of the above syntypes should be designated as lectotype it would result in a name
change for the affected species, assuming that the western forms of one or both of these species
are indistinguishable from those to the east. Inasmuch as systematic studies on these species have
not yet been completed, I consider it best to defer action on the problem at this time. Original
description date 25 April 1857.
= a senior synonym of Notropis stramineus (Cope) and Notropis colucellus (Cope)
Cyprinella lugubris Girard 1857
Girard 1857:199. TYPE LOCALITY: Cottonwood R., ca. 5 mi. NW of Durham, Marion Co.,
Kansas. HOLOTYPE: USNM 141 (one set of pharyngeal arches in bone collection), F. Creutzfeldt,
34 July 1853.
REMARKS.-For discussion of correct type locality see REMARKS under Cyprinella gunnisoni.
Girard (1858:271) listed one type, which is thus the holotype. Counts: teeth 2,4-4.1, with well-
developed hooks; no serrations; the lesser teeth long and slender. In addition to the above tooth
count, Girard (op. cit.: 271) listed 9 or 10 anal rays (his method of listing counts is difficult to in-
terpret) and an elongate, subfusiform body. Based on these counts and observations and Cross's
(1967) range maps, C. lugubris most likely is a synonym of N. rubellus, with N. pilshryi the next
best possibility. C. lugubris was erroneously synonymized by Jordan and Gilbert (1883:177) with
Cyprinella rnacrostoima (= Notropis amnabilis). Original description date 25 April 1857.
= most likely Notropis rubellus (Agassiz), but possibly Notropis pilsbryi Fowler
Hydrophlox lutipinnis Jordan and Bravton 1878
Jordan and Bravton 1878:36-37. TYPE LOCALITY: headwaters of Oconee R. (at Sulphur Springs
and Fuller's Mills), Hall Co., Georgia. SYNTYPES: USNM 31090 (6, 43.1-55.8), MCZ 24386 (1,
37.0), CAS-SU 25(0 (1, 58.5), BMNH 1880.1.21.3 (1, 38.0), D. S. Jordan and A. W. Bravton, sum-
mer 1877.
REMARKS.-Counts: USNM 31090 (teeth 2,4-4,2 in one: anal rays 8 in six); MCZ 24386 (anal
rays 8). Original description date (31 December) 1878.
= Notropis lutipinnis (Jordan and Brayton)
Leuciscus lutrensis Baird and Girard 1853
Baird and Girard 1853:391. TYPE LOCALITY: Otter Cr., trib. to N. Fk. of Red R., either Kiowa
or Tillman Co., Oklahoma. (Locality erroneously given in original description as being in Arkan-
sas). SYNTYPE: USNM 104 (1, 44.8), R. B. Marcv and G. B. McClellan, 1852.
REMARKs.-Girard (1858:272-273) listed five original syntypes (USNM 104). He also listed 10
specimens (USNM 107) from Gypsum Creek, a tributary of the Canadian River, but these were
not collected till late August 1853 (Whipple 1854:18-19; 1856: appendix D), and thus almost
certainly could not have been examined by Baird and Girard when preparing the original descrip-
tion. leuciscus bubalinus, which was originally described in the same paper as Lecciscus lutren-
sis, was found by Hubbs and Ortenburger (1929a:34) to be identical with that species. As first
revisers they selected the name lutrensis over bubalinus. Contreras (1975) reviewed species in
Mexico and the Rio Grande basin. Counts for extant syntype: teeth 4-4; anal rays 9. Original
description date 5 September 1853.
= Notropis lutrensis lutrensis (Baird and Girard)
Cyprinella luxiloides Girard 1857
Girard 1857:198. TYPE LOCALITY: San Pedro Cr., trib. to Rio San Antonio, Texas. TYPES: None
located. Girard (1859a:55) listed two specimens (USNM 131), collected by Dr. C. B. Kennerly in
1854.
REMARKs.-Although this species cannot be positively identified, it most likely is a synonym
of N. aimahilis (Girard). Jordan and Gilbert (1883:177) so indicated this when they synonymized
1978
BULLETIN FLORIDA STATE MUSEUM
C. luxiloides with Cyprinella macrostoma (=Notropis armahilis). Hubbs, Kuehne, and Ball
(1953:226), however, regarded luxiloides as a subspecies of N. lutrensis. Girard's (1859a: pl. 31,
fig. 13) illustration of C. luxiloides is very similar to those of C. macrostoma (op. cit.: pl. 31, fig. 5)
and Alburnellus amabilis (op. cit.: pl. 29, fig. 10), particularly with regard to scale size (only five
visible body-circumference scales above lateral line), number of anal rays (nine), large eye size,
relative positions of the fins, and pigmentation pattern. The only other species of Notropis in the
Rio San Antonio drainage to which the above characters could possibly apply is N. tolurellus,
but the number of anal rays in that species is almost always eight. Original description date 25
April 1857.
= probably Notropis amabilis (Girard)
Notropis lythrurus Jordan 1884
Jordan, in Jordan and Meek 1884:476. TYPE LOCALITY: Grand River (or tributary thereof),
Lansing, Ingham Co., Michigan. SYNTYPES: Possibly ANSP 3054-3055 (2, 49.0-50.0). These speci-
mens bear only the data "Michigan, E. D. Cope." and there is no positive indication that they
actually are types.
REMARKs.-The name lythrurus was first proposed in a footnote (Jordan and Meek 1884:476),
as follows: "Notropis lithrurus, Jordan, nom. sp. nov. = Minnilus diplaemius, Jordan and Gilbert,
Synopsis Fish. N. Am., p. 197= Hypsilepis diplaemius, Cope, not Semotilus diplaemirns, Rafi-
nesque." The incorrect spelling was emended to lythrurus in the "List of Corrections" on intro-
ductory page VIII in the original volume. Although this does not, by itself, constitute a descrip-
tion, the new species name is available (according to Article 16a, International Code of Zoologi-
cal Nomenclature) as a substitute name for Cope's Hypsilepis diplaemius, which is a species
presumably different from Rafinesque's Senotilus diplemia. Cope's (1868a:162-163) account
stands as the original description of N. lythrurus, which in turn is a junior synonym of N. um-
bratilis cyanocephalus (Copeland 1877). Jordan and Evermann's (1896:300) designations of White
River, Indianapolis, Indiana as the type locality and USNM 20115 as the type specimen of N.
lythrurus are invalid. Snelson and Pflieger (1975:235-236) also have discussed this situation. Orig-
inal description date 23 October 1884.
= Notropis umbratilis cyanocephalus (Copeland)
Notropis ,macdonaldi Jordan and Jenkins 1889
Jordan and Jenkins, in Jordan 1889a:354; pl. 44, fig. 4. TYPE LOCALITY: Shenandoah River,
at Waynesboro, Augusta Co., Virginia; North River, Loch Laird, Virginia; and Buffalo Creek,
near Lexington, Virginia. Type locality here restricted to Shenandoah River. LECTOTYPE: USNM
39859 (93.0), D. S. Jordan and C. H. Gilbert, summer 1888. Lectotype herein designated. LECTO-
PARATYPES: USNM 214481 (1), paratopotype; CAS 29303 (old IU 7817) (18, plus one specimen of
N. procne). CAS-SU 597 (1), which Bohlke (1953:35) listed as a "paratype," is not so regarded
here, because the listed locality (Luray, Virginia) was not one of the localities cited in the original
description.
REMARKs.-H-ubbs and Moore (1940:96) were first to identify N. macdonaldi as a hybrid.
Original description date 5 July 1889.
= Hybrid: Notropis corn utus x Notropis rubellus
Notropis lmacrolepidotus Forbes 1885
Forbes 1885:138. TYPE LOCALITY: "Illinois" (no precise locality given). TYPES: None located.
Forbes listed one specimen in original description.
REMARKS.-Snelson (1973) reviewed the status of this form. The original description could
apply either to N. fumes or N. umbratilis cyanocephalus, but the high predorsal scale count (19)
is outside the range of either species. Forbes (1884:76) first introduced the name macrolepidotus
as a nnmen nudum, and stated that the species was rare and known only from the Illinois River.
N. funieus is not known today from that river system (although N. umbratilis does occur there),
but conceivably could have lived there in the past. Original description date (31) March 1885.
Vol. 23, No. 1
1978
GILBERT: NOTROPIS TYPE CATALOGUE
=either Notropis umnbratilis cyanocephalus (Copeland) or a senior svnonvni of Notropis
fuimeus Evermann
Cyprinella macrostoma Girard 1857
Girard 1857:198. TYPE LOCALITY: Devils River, probably just above month into Rio Grande
(or possibly ca 45 mi upstream), Val Verde Co., Texas: and Rio San Juan, near China, Nuevo
Leon, Mexico. SYNTYPES: USNM 129 (3, 36.6-48.0) (from Devils River), John H. Clark, 1851;
USNM 130 (one right pharyngeal arch only) and MCZ 1798 (ex USNM 130) (1,45.0) (from China),
Lt. D. N. Couch, 1853.
REMARKs.-Girard (1859a:54) listed three original syntypes for USNM 130. Although Cypri-
nella macrostomna was generally considered to be a valid species for many years following its
original description, the last literature reference appears to be that by Jordan, Evermann, and
Clark (1930:131). The types have been examined independently by Carl L. Hubbs, Robert R.
Miller, and myself, and all of us agree that they represent the species called X. amabilis, which
was described by Girard (1857:193) in the same paper. Miller (1976:11), as first reviser, formally
selected the name amiabilis, thus relegating C. macrostomla to the synonymy of that species. Al-
though Contreras (1967:6-7) did not at first indicate that N. oamahilis occurred in the state of
Nuevo Leon, he later (Contreras 1974:192) listed it among those species that apparently have
become extirpated from the area. This lends credence to the provenance of Couch's material,
which had formerly been in doubt. Counts for MCZ 1798: teeth 2,4-4,2; anal rays 10. Counts for
USNM 129: teeth 2,4- in largest specimen; anal rays 9 (1), 10 (2). Original description date 25
April 1857.
= Notropis amabilis (Girard)
Hemitremia maculata Hay 1881
Hay 1881:505-506. TYPE LOCALITY: Chickasawha R. (and tributaries), near Enterprise, Clarke
Co., Mississippi. SYNTYPES: USNM 27438 (3, 27.7-30.5) (= N. rolucellus), USNM 101174 (1, 45.3)
(=N. maculatus), O. P. Hay, March-April 1880. Jordan and Evermann (1896:259) also listed
USNM 32245 as comprising a syntypic series, but since neither this catalogue number nor the
locality from which the series came (Jackson, Mississippi) were included by Hay in his original
description, Jordan and Evermann's action is considered to be erroneous.
REMARKS.-The extant types of this species are in extremely poor condition. The three syn-
types in USNM 27438 were identified as N. rolucellus bv R. D. Suttkus on 23 June 1954, a diag-
nosis with which I concur. The other specimen (USNM 101174), which is erroneously labelled
a "paratype," was catalogued (without explanation) into the USNM collection on 9 November
1935. Although nothing indicates that it was removed from USNM 27438, it is considered a valid
type because of its condition (similar to that of the other three svntypes) and the presence of
Hay's original label in the jar. The main diagnostic character of N. maculatus (a well developed
caudal spot) is not evident in this specimen, but this is believed to be the result of the caudal fin
having been broken off anterior to where the spot occurs. Both pharyngeal arches are missing
(they were not located in the osteological collection), and the anal rays, though difficult to count,
appear to be eight in number (the count listed by Hay). There are about 20 bodv-circumference
scales and about 14 predorsal scales (Hay listed 16 predorsal scales in the original description).
Perhaps most importantly, the mouth is small and moderately oblique, with its posterior margin
falling well short of the eye. The melanophores in the region of the lateral stripe are of the same
distribution and intensity as in specimens examined of N. mnaculatus. Despite its poor condition, I
identify this specimen as N. nmaculatus (as presently recognized), and recommend that it be
chosen as lectotype of Hemitremia miaulata. Original description date 23 February 1881.
= Notropis maculatus (Hay)
Alburnellus matutinus Cope 1870
Cope 1870:465. TYPE LOCALITY: Neuse R., Wake Co., North Carolina. SYNTYPES: Two svn-
types (ANSP 3844-3845) are listed in the ANSP catalog, but these cannot be located.
REMARKs.--.. matutinus was until recently considered a valid species. Recent unpublished
BULLETIN FLORIDA STATE MUSEUM
Vol. 23, No. 1
studies by F. F. Snelson, Jr. indicate that this form is not even subspecifically distinct from N.
gardens. Original description date 21 November 1870.
= Notrropis gardens (Cope)
Notropis mnearnsi Snyder 1915
Snyder 1915:582-584, fig. TYPE LOCALITY: San Bernardino R., trib. to Yaqui R., Sonora, Mex-
ico (near international boundary monument 77). HOLOTYPE: USNM 76163 (39.2), Dr. E. A.
Mearns, 6 October 1893. PARATYPES: USNM 76164 (4), paratopotypes.
REMARKS.-Miller and Simon (1943) regarded this as a questionably valid species, and Carl
Hubbs (1954:293) and Minckley (1973:138-139) called it a subspecies of N. fornosus. It has usually
been considered a junior synonym of N. formosus (Girard), which Contreras (1975), in turn, now
regards as a subspecies of N. lutrensis (Baird and Girard). Counts: teeth 4-4 in holotype and para-
types. Original description date 23 December 1915.
= Notropis lutrensis fornosus (Girard)
tNotropis megalepis Smith 1962
Smith 1962:511-512, fig. 4. TYPE LOCALITY: Ogallala Formation, marl deposit located in Sec.
7, T.11S, R.37W, Logan Co., Kansas. (Pliocene age). HOLOTYPE: KU F849, a complete fossil speci-
men, 91 mm SL, Claude W. Hibbard et al.
REMARKS.-This is the only fossil cyprinid species currently assigned to Notropis. In the origi-
nal description, Smith stated that the generic assignment was tentative and was based on the
large scales, general shape and appearance of the body, and absence of other distinctive charac-
ters. Original description date 28 September 1962.
= tNotropis megalepis Smith
Cyprinus miegalops Rafinesque 1817
Rafinesque 1817:121. TYPE LOCALITY: Hudson R. system, New York. TYPES: Not located.
REMARKS.-The name mnegalops was not used for many years after its original proposal, until
Jordan (1885a:123) resurrected it as a senior synonym of Cyprinus cornmtus Mitchill, on the
grounds that Mitchill's (1817:289) preliminary account of that species did not constitute an ade-
quate description. Jordan and Evermann (1896:281-282) subsequently reversed this decision,
and the name cornitus has been used since (see Jordan 1916:28, and Gilbert 1964:142-143). Orig-
inal description date (31) December 1817.
= Notropis cornutus (Mitchill)
Alburnus megalops Girard 1857
Girard 1857:193. TYPE LOCALITY: San Felipe Cr., near Del Rio, Val Verde Co., Texas. SYN-
TYPES: USNM 36529 (ex USNM 69) (1, 40.2), ANSP 3246 (ex USNM 69) (1, 36.5), MCZ 1682 (ex
USNM 69) (2, 35.0-39.5), John H. Clark, 1851. (Label in jar of MCZ types says "1853").
REMARKs.-Girard (1859a: pl. 29, figs. 1-4) illustrated one of the syntypes and listed 20 speci-
mens in the original type series (USNM 69) (op. cit.: 52). Fowler (1910: pl. 20, fig. 44) illustrated
the ANSP "cotype." Jordan (1885a:123) substituted the species name scwaini for megalops, which
is preoccupied in Notropis by Cyprinus megalops Rafinesque 1817. Hubbs, Kuehne, and Ball
(1953:226) indicated that specimens recorded as N. swaini by Jordan and Gilbert (1886:24) and
Evermann (1892:79) from the Guadelupe River system are N. amahilis. This apparently was the
first time that megalops was synonymized with amahilis (both were described in the same paper)
but as they said nothing about the relationships of these two forms throughout their overall
ranges, Hubbs, Kuehne, and Ball cannot properly be regarded as first revisers. For further dis-
cussion see account of Alburnus amabilis.
Counts for ANSP 3246: teeth 2,4-4,2; anal rays 9. Counts for MCZ 1682: teeth 2,4-4,2; pre-
dorsal scales 16; body-circumference scales 13-2-9 (larger specimen); teeth -4,2; anal rays 10;
predorsal scales 15; body-circumference scales 11-2-10 (smaller specimen).
Original description date 25 April 1857.
= Notropis amabilis (Girard)
GILBERT: NOTROPIS TYPE CATALOGUE
Trycherodon megalops Forbes 1883
Forbes, in Jordan and Gilbert 1883:247. TYPE LOCALITY: Illinois R. (Long Lake), Pekin, Illi-
nois. LECTOTYPE: USNM 28406 (40.4), S. A. Forbes, 27 October 1879. Gilbert and Bailey (1972:21)
designated lectotype. LECTOPARATYPE: GAS 13513 (old IU 4734) (1).
REMARKS.-Jordan (1884b:821) was first to synonymize Trycherodon megalops with Opso-
poeodus emiliae. Schrenkeisen (1938:107) and Hubbs and Lagler (1947:64-65) subsequently re-
garded it as the northern subspecies of Opsopoeodus emiliae. Gilbert and Bailey (1972:13) again
relegated megalops to the synonymy of emiliae. Original description date 1 April 1883.
= Notropis emiliae emiliae (Hay)
Notropis mekistocholas Snelson 1971
Snelson 1971:449-462, fig. la-b. TYPE LOCALITY: Rocky R., at st. hwy. 902 bridge, 7.0 air mi
SW center of Pittsboro, Chatham Co., North Carolina. HOLOTYPE: USNM 205299 (ex NCSM
4928) (55.7), tuberculate adult male, F. F. Snelson, Jr. and W. M. Palmer, 2 June 1968 (orig. no.
FFS 68-31). PARATYPES: USNM 2053() (8), UMMZ 197680 (7), UF 17303 (7), CU 64397 (7), TU
63272 (7), paratopotvpes.
REMARKS.-Original description date 3 September 1971.
= Notropis mekistocholas Snelson
Cyprinus melanturus Rafinesque 1817
Rafinesque 1817:121. TYPE LO(:ALITY: Hudson R. system, New York. TYPES: None located.
REMARKS.-Jordan (1877d:12) was first to synonvmize this form with N. cornutus. Original
description date (31) December 1817.
= Notropis cornutus (Mitchill)
Notropis metallicus Jordan and Meek 1884
Jordan and Meek 1884:475-476. TYPE L)OALITY: Alapaha R., tributary to Suwannee R., at
Nashville, Berrien Co., Georgia. This river was erroneously said to be the Altamaha in the original
description. SYNTYPES: USNM 28511 (6, 22.5-34.2), CAS-SU 2527 (1, 29.0), W. J. Taylor.
REMARKS.-Fowler (1941:234) seems to be the first to have svnonvmized this form with V.
hypselopterus. Original description date 23 October 1884.
= Notropis hypselopterus hypselopterus (Giinther)
Aztecula moexicana Meek 1902
Meek 1902:81-82; pl. 18, upper fig. TYPE LOCALITY: Rio Moctezuma, San Juan del Rio, Que-
retaro, Mexico. HOLOTYPE: FMNH 3606 (58.6), S. E. Meek and Frank Eugene Lutz, 16 May 1901.
PARATYPES: FMNH 80452 (90), UMMZ 162462 (ex FMNH 80452) (10), paratopotypes.
REMARKS.-Contreras (1974) listed Notropis mexicanus as a valid species, but Miller (1976:10)
tentatively synonymized it with N. sallci. Original description date (31) May 1902.
= Notropis sailei (Giinther)
Alburnellus micropteryx Cope 1868
Cope 1868b:233. TYPE LOCALITY: Holston R., Virginia. LECTOTYPE: ANSP 2842 (54.9), E. D.
Cope, October 1867. Fowler (1910: pl. 21, fig. 54) designated lectotype by illustrating "type"
(ANSP 2842); calculated length 57.7 mm SL. LECTOPARATYPE: ANSP 2843 (1), paratopotype.
REMARKS.-Condition of types fair to poor for old specimens (fins broken, but characters
recognizable). Counts: teeth 2,4-4,2 (lectotype), 1,4-4,2 (lectoparatype); anal rays 10 (lectotype),
9 (lectoparatype). This form was long called a valid species (Kuhne 1939:49), but more recently
has been called a subspecies of N. rubellus (Smith-Vaniz 1968:49) or synonymized with that spe-
cies (Moore 1957: 125). The systematics of the N. robellus complex have not yet been adequately
studied. Original description date 2 December 1868.
= Notropis rubellus (Agassiz)
Minnilus microstomus Rafinesque 1820
Rafinesque 1820b:47. TYPE LOCALITY: Kentucky R., Kentucky. TYPES: None located.
BULLETIN FLORIDA STATE MUSEUM
REMARKS.-This form cannot be identified with certainty. Jordan (1876a:94; 1876b:282) con-
sidered it to be a valid species, and initially synonymized Hyhopsis gracilis Agassiz and Hybopsis
longiceps Cope with it. Later, he (Jordan 1885b:811) wrote, "The identification of Rafinesque's
Minnilus microstonus is too uncertain to warrant the use of his name." M. microstomus was said
to be a small species with a straight lateral line, olivaceous back and head, a few black dots on
the sides, a small mouth, and with eight rays in both the dorsal and anal fins. This description
fits N. stramineus or N. colucellus as well as any species known from northern Kentucky. Original
description date (31) May 1820.
= identification uncertain. Possibly a senior synonym of either Notropis stramiinus (Cope) or
Notropis volucellus (Cope)
Hyhopsis missuriensis Cope 1871
Cope 1871:437-438. TYPE LOCALITY: tributary to Missouri R., near St. Joseph, Missouri. LEC-
TOTYPE: ANSP 4374 (40.0), Dr. William Stimpson. Fowler (1910: pl. 15, fig. 6) designated lecto-
type by illustrating "type" (ANSP 4374); calculated length 40.2 mm SL. LECTOPARATYPES: ANSP
4375-4377 (3), paratopotypes.
REMARKS.-Counts for original syntypes: teeth -4 (1); anal rays 7 (4); predorsal scales 13 (1),
14 (2), 15 (1). Hubbs and Greene (1928:377) synonymized Hyhopsis scylla and H. missuriensis,
which were both described in the same paper; as first revisers, they chose the latter name. Bailey
and Allum (1962:64-68) showed this form to be a valid subspecies, although the type locality lies
in the zone of intergradation between the subspecies stramineus and missuriensis. Original de-
scription date (31 December) 1871.
= Notropis stramineus missuriensis (Cope)
Hyhopsis montanus Meek 1885
Meek 1885:526-527. TYPE LOCALITY: upper Missouri River basin (no further data). SYNTYPES:
USNM 36882 (3, 46.4-54.6), Dr. F. V. Hayden.
REMARKS.-Syntypes in fairly good condition for old specimens. Counts: teeth 1,4-4,1 (2),
-4,1 (1); anal rays 8 in all (fin in one specimen with one of middle rays missing); predorsal scales
absent (2) or present over posterior third of area (1). Bailey and Allum (1962:68) erroneously
synonymized this form with N. stramineus missuriensis. Original description date 25 February
1885.
= Notropis dorsalis (Agassiz)
Cliola montiregis Cope 1885
Cope 1885:168. TYPE LOCALITY: Monterrey, Nuevo Leon, Mexico. HOLOTYPE: ANSP 19344
(37.5), E. D. Cope, November 1883.
REMARKs.-The type of this species is apparently unique and thus is considered the holotype.
Fowler (1910: pl. 17, fig. 21) illustrated "type;" calculated length 38.1 mm SL. Counts for holo-
type: teeth 1,4-4,0; anal rays 8. Jordan and Evermann (1896:272) were first to synonymize this
form with N. lutrensis. Original description date 7 March 1885.
= Notropis lutrensis lutrensis (Baird and Girard)
Notropis moorlesi DeBuen 1956
DeBuen 1956:527-541, fig. 1. TYPE LOCALITY: dentro del Rio Tepelneme, tributary to Rio
Santo Domingo (tributary of Rio Papaloapan), Presa de El Capulin, near pueblo de Tepelneme,
Oaxaca, Mexico. HOLOTYPE: Instituto de Biologia (Mexico) uncat. (?) (60.0), Salvador Morales
and Dr. Fernando deBuen, 24 February 1955. PARATYPES: UMMZ 172459 (13); remaining 20
paratypes divided among la Comision del Papaloapan, la Comision de Piscicultura Rural de la
Direccion General de Pesca e Industrias Conexas, and el Instituto Politecnico.
REMARKS.-This species frequently has a barbel at each corner of the mouth. Original descrip-
tion date 20 June 1956.
= Notropis moralesi deBuen
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
Notropis muskoka Meek 1899
Meek 1899:308. TYPE LOCALITY: lower part of Cull Lake, near Gravenhurst, Ontario, Canada.
HOLOTYPE: FMNH 2964 (59.5), S. E. Meek, mid-September 1899. PAHATYPES: FMNH 2964a (20),
USNM 126873 (ex FMNH 2964a) (2), CAS-SU 21483 (ex FMNH 2964a) (4), paratopotypes.
REMARKS.-Huhbs (1926:41) was first to svnonymize this form with N. heterolepis. Original
description date (30) November 1899.
= Notropis heterolepis Eigenmann and Eigenmann
Nototropis nazas Meek 1904
Meek 1904: 70, fig. 19. TYPE LOCALITY: headwaters of Rio Nazas, Santiago Papasquiaro,
Durango, Mexico. HOLOTYPE: FMNH 4375 (49.8), S. E. Meek, 25 May 1903. PARATYPES: FMNH
33572-33704 (133), paratopotypes.
REMARKs.-Original description date (31) August 1904.
= .otropis nazas (Meek)
Notropis altipinnis neusensis Hubbs and Raney 1948
Hnbbs and Raney 1948:8-10, pl. 1. TYPE LOCALITY: trib. of Trent R., 7 mi. VWSW of New Bern,
Craven Co., North Carolina. HOLOTYPE: UMMZ 136152 (43.7), E. C. Raney, E. A. Lachner, and
R. A. Pfeiffer, 30 March 1941. PARATYPES: UMMZ 136153 (4), paratopotypes; UMMZ 136154
(6), CU 9658 (2), CU 10579 (58 originally, now 56).
REMARKS.-This form and others described in the same paper are not generally regarded as
valid, though they have never been formally synonymized. Original description date 26 April
1948.
= Notropis altipinnis (Cope)
Minnilus nigripinnis Gilbert 1884
Gilbert 1884c:14. TYPE LO(CALITY: Shunganunga Cr. tribe to Kansas R.), near Topeka, Kansas.
HOLOTYPE: USNM 36613 (59.7), F. W. Cragin, 1884. PARATYPES: USNM 36608 (2), paratopo-
types. (Only one paratype is mentioned in original description.)
REMARKs.-Gilbert (1885:513-514) subsequently republished original description verbatim
"for convenience of reference." Jordan (1885b:814) was first to svnonvmize this form with N.
umbratilis. Snelson and Pflieger (1975:236) discussed the status of this form. Original description
date (30) September 1884.
= Notropis umbratilis iumbratilis (Girard)
Graodus nigrotaeniatus Giinther 1868
Giinther 1868:485-486. TYPE LOCALITY: Atlisco, Mexico. SYNTYPES: BMNH 1868.3.3.19-21
(3, 68.5-76.0), M. Boucard.
REMARKs.-Jordan (1880a:226) discussed the syntypes and suggested that the unusual denti-
tion represents an aberrant condition. Meek (1904:68) was first to synonymize this form with
N. boucardi (based on observations of C. Tate Regan, who examined the types of both), and as
first reviser, selected the latter name. Original description date 14 March 1868.
= Notropis boucardi (Giinther)
Alburnus nitidus Kirtland 1854
Kirtland 1854:4445, fig. TYPE LOCALITY: Rocky River, near Cleveland, Ohio. TYPES: None
located. The original description mentioned an undetermined number of syntypes, collected by
J. P. Kirtland on 9-12 October 1853, as having been presented to Cleveland Academy of Science
at meeting of 21 March 1854.
REMARKS.-Jordan and Gilbert (1883:202) erroneously synonymized this form with Minnilus
(= Votropis) rubellus. Jordan and Evermann (1896:293) were first to synonymize it with N.
atherinoides. Original description date 1 February 1854.
= Notropis atherinoides Rafinesque
BULLETIN FLORIDA STATE MUSEUM
Moniana nitida Girard 1857
Girard 1857:201. TYPE LOCALITY: Rio Monterrey trial to Rio San Juan), Cadereita (=Ca-
dereyta), Nuevo Leon, Mexico. SYNTYPES: USNM 96 (one set of pharyngeal teeth in bone collec-
tion), USNM 39657 (2, 47.5-52.4), Lt. D. N. Couch, winter 1852-1853.
REMAHKS.-Girard (1859a:58) listed only two original syntypes. Possibly the set of pharyngeal
arches (USNM 96) was removed from one of the specimens now comprising USNM 39657, inas-
much as the arches are missing from one of the two syntypes. Counts for USNM 96: teeth 1,4-
4,0. Counts for USNM 39657: teeth 4-4 (1); anal rays 7 in both. Jordan and Gilbert (1883:175)
erroneously synonymized this form with Cliola deliciosa (=N. stramineus), but it was later re-
validated by Jordan (1885b:811). Jordan and Evermann (1896:264-265) pointed out that Moniana
nitida is preoccupied in Notropis by Alburnus nitidus Kirtland 1854, and therefore proposed N.
braytoni as a substitute name for Girard's species. Original description date 25 April 1857.
= a preoccupied senior synonym of Notropis braytoni Jordan and Evermann
Hybopsis niveus Cope 1870
Cope 1870:460-461. TYPE LOCALITY: Upper waters of Catawba R., North Carolina. LECTO-
TYPE: ANSP 2930 (51.0), E. D. Cope, Autumn 1869. Fowler (1910: pl. 18, fig. 27) designated
lectotype by illustrating "type" (ANSP 2930); calculated length 52.7 mm SL. LECTOPARATYPES:
ANSP 2931-2947 (17), paratopotypes.
REMARKS.-Gibbs (1955) reviewed species. Condition of types fair to poor. Fowler (1910:282)
listed 19 syntypes, but only 17 were subsequently found. Counts for largest syntype: teeth 4-4;
anal rays 8. Original description date 21 November 1870.
= Notropis niveus (Cope)
Notropis nocomis Jordan and Gilbert 1886
Jordan and Gilbert 1886:23-24. TYPE LOCALITY: Rio Comal, at New Braunfels, Comal Co.,
Texas. LECTOTYPE: USNM 36531 (39.6), D. S. Jordan, C. H. Gilbert, J. Swain, and S. E. Meek;
July, August, or September 1884. Hubbs (1951a:11-12) designated lectotype. LECTOPARATYPES:
USNM 93516 (55), paratopotypes; USNM 36514 (9), USNM 46219 (11).
REMARKs.-This name first appeared (Jordan 1885b:812) as a nomen ndumn. It was subse-
quently suppressed by Jordan and Gilbert (1886:23-24), who regarded it as identical to N. delici-
osuis (= N. straminens), but at the same time they gave diagnostic characters for their rejected
species. The name nocomis could have remained without standing, according to the Interna-
tional Code of Zoological Nomenclature (1964: Article lid), but Hubbs (1951a) elected to resur-
rect it as a subspecies of N. colucellus. This action served to make N. noconis Evermann 1892 a
junior primary subjective homonyn, for which the name N. amnis Hubbs and Greene, in Hubbs
1951, was substituted. The nomenclatural origin of N. nocomis Jordan and Gilbert closely paral-
lels that of N. ionthas, a name that was similarly rejected in the same paper (Jordan and Gilbert
1886:11), but which has since been found to represent a valid species. Unlike nocomis, however,
the name ionthas still has no legal nomenclatural standing (see Snelson and Jenkins 1973:293).
Although N. colmuellus nocomis is not indicated below as being a valid subspecies, subsequent
study may prove it to be so.
Original description date 17 September 1886.
= Notropis colucellus (Cope)
Notropis nocomis Evermann 1892
Evermann 1892:78-79. TYPE LOCALITY: Trinity R., Magnolia Point, 10 mi SW of Palestine, An-
derson Co., Texas. LECTOTYPE: USNM 45556 (45.0), B. W. Evermann, R. R. Gurley, and J. A.
Single, 23 November 1891. Evermann and Kendall (1894: pl. 17) designated lectotype by illus-
trating "type." This specimen is also the holotype of N. amnis pinnosa Hubbs and Bonham. LEC-
TOPARATYPES: USNM 93517 (20), USNM 125175 (apparently ex USNM 93517) (21), CAS-SU
21146 (apparently ex USNM 93517) (24), CAS 13517 (old IU 9041) (4), paratopotypes. Other
syntypes (USNM 58792) were identified by Hubbs (1951a:12) as Dionda episcopal couchi.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
REMARKS.-This form is a junior homonym of N. nocmnis Jordan and Gilbert (1886), as a re-
sult of Hubbs' (1951a) action revalidating the latter species (see preceding account). It also is a
senior synonym of N. amnis pinnosa, but Clemmer (1970) found that subspecies not to be valid.
Original description date 25 May 1892.
= a preoccupied senior synonym of Notropis amnis Hubbs and Greene
Notropis norrisi Fowler 1945
Fowler 1945:235-236, figs. 114 and 116. TYPE LOCALITY: basin of Altamaha R., near Fitz-
gerald, Georgia. (This locality very likely is Osewitchee Springs, 14.5 mi. N of Fitzgerald, Wilcox
County.) HOLOTYPE: ANSP 70705 (39.0), Robert Norris, 1941. PARATYPES: ANSP 70706-70710
(5), paratopotypes.
REMARKs.-Suttkus and Raney (1955a:84) were first to synonymize this form with Hybopsis
harper (= N. harperi. Contrary to their opinion that the types of N. norrisi are probably from
the Suwannee River system, they are more likely from the Altamaha system, as stated in the
original description. Original description date 25 October 1945.
= Notropis harper Fowler
Cyprinella notata Girard 1857
Girard 1857:198. TYPE LOCALITY: Rio Seco (= Seco Creek), at D'Hanis, Medina Co., Texas.
LECTOTYPE: USNM 136 (38.5), Dr. C. B. Kennerly, 1853. Gibbs (1957:180-181) designated lecto-
type. Despite statement by Clark Hubbs (1954b:280), Jordan (1885a:124) did not designate lec-
totype for this species. LECTOPARATYPE: USNM 163955 (1). Third syntype is N. lutrensis and is
recatalogued under that name.
REMARKS.-Girard (1858:270: pl. 58, figs. 16-20) listed only two original syntypes and illus-
trated one of these. Moore (1952) used the specific name notatus for the blacktail shiner, based
on the then current nomenclatural rule of line priority. Clark Hubbs (1954b:280) was first to
synonymize this form with N. cenustus, and, as first reviser, selected latter name. Original
description date 25 April 1857.
= Notropis renuistus renustus (Girard)
Notropis notemiigonoides Evermann 1892
Evermann 1892:81-82. TYPE LOCALITY: Neches R., ca 14 mi E of Palestine, Anderson County,
Texas (at Palestine and Rusk RR bridge). LECTOTYPE: USNM 125183 (50.5), B. W. Evermann,
J. T. Scovell, R. R. Gurley, and J. A. Single, 24 November 1891. Evermann and Kendall (1894: pl.
18) designated lectotype, by illustrating "type." This antedates Jordan and Evermann's listing
of USNM 45559 as the "type." LECTOPARATYPE: USNM 203330 (1). The other four syntypes are
all N. fumers; one was taken with the above specimens of N. umbratilis, and the other three were
collected from Sims Bayou, near Houston, Texas.
REMARKs.-Ortenburger and Hubbs (1927:131) pointed out the close similarity of N. note-
migonoides to N. furmeus and N. umhratilis, but did not synonymize it with either species. Snel-
son (1973:169-170) and Snelson and Pflieger (1975:236) discussed the status of N. notemigonoides
and gave reasons for synonymizing it with N. umbratilis rather than N. fumeus. Original descrip-
tion date 25 May 1892.
= Notropis umhbratilis cynocephalus (Copeland)
Alburnops nubilus Forbes 1878
Forbes, in Jordan 1878b:56-57. TYPE LOCALITY: Rock R., at Oregon, Ogle County, Illinois.
SYNTYPES: INHS 26945 (6, 46.6-55.3), Stephen A. Forbes, 6 May 1877. Jordan and Evermann
listed USNM 28410 as "type," but I have not located this series. Bohlke (1953:30) erroneously
considered this to constitute a lectotype designation. Bohlke (1953) also listed SU 1280 (= CAS-
SU 1280) as "paratypes" (actually syntypes), but the three specimens involved are from Belvidere,
Illinois, a locality not included in Forbes' original description.
BULLETIN FLORIDA STATE MUSEUM
REMARKS: N. nubilus was long assigned to the genus Dionda, based on a long gut tract. Swift
(1970:3081-B) was the first to place it in Notropis. Original description date (30) June 1878.
= Notropis nubilus (Forbes)
Notropis nux Evermann 1892
Evermann 1892:77-78. TYPE LOCALITY: Trinity R., at Magnolia Point, 10 mi SW of Palestine,
Anderson County, Texas. See REMARKS. LECTOTYPE: USNM 45555 (45.6), B. W. Evermann, J. T.
Scovell, and R. R. Gurley, 23 November 1891. Evermann and Kendall (1894: pl. 17, top fig.)
designated lectotype by illustrating "type." See REMARKS. LECTOPARATYPES: USNM 125166 (10)
(from Neches R., near Palestine, Texas). A single paratype from Long Lake, also near Palestine,
Texas, cannot be located.
REMARKS.-The correct type locality and source of the lectotype designation for N. nux are
confused. Although Suttkus (1958:308) designated a lectotype (USNM 45555), his action was
superceded by that of Evermann and Kendall (1894: pl. 17, top fig.), whose illustration of the
"type" fulfills the requirements for a lectotype designation, as enumerated in Article 74b of the
International Code. According to Evermann and Kendall (1894), the specimen illustrated is from
the "Neches River, Palestine, Texas," whereas the specimen designated by Suttkus (1958) is from
the Trinity River, also near Palestine, Texas. This would suggest that the individual illustrated
is from the series now catalogued as USNM 125166 (from the Neches R.), but Jordan and Ever-
mann (1900:3241, pl. 47, fig. 117) gave the locality for the same figure appearing in Evermann
and Kendall (1894) as the Trinity River. Furthermore, the following information appears in the
fish catalogue with the entry for USNM 45555: "Notropis nux (type) Trinity River, Palestine,
Texas. November 23, 1891. alc. Recd. from U. S. Fish Comm. collected by B. W. Evermann.
Catalogued Oct. 26, 1894. 1 spec. 'Drawn'." An old label in the jar itself contains the above lo-
cality information, together with the notation "Specimen drawn." Although there is neither a
scale of measurement nor other indication of length of the illustration in plate 17, examination
of the specimen catalogued as USNM 45555 and comparison with the drawing indicate that it
is the one Evermann and Kendall illustrated, despite the discrepancy in locality data. Finally,
the Trinity River specimen (no. 45555) is larger and in better condition than any of those in the
other series. Based on the above, it seems plausible to assume that the locality data given by
Evermann and Kendall (1894: pl. 17) is in error, the Trinity River is the correct type locality, and
that the lectotype designations by Evermann and Kendall and by Suttkus are based on the same
specimen. Bailey, Winn, and Smith (1954:126) were first to synonymize N. nux with N. roses
(= N. texanus). Original description date 25 May 1892.
= Notropis texanus (Girard)
Leuciscus obesus Storer 1845
Storer 1845:48. TYPE LOCALITY: Florence, Alabama. TYPES: None located.
REMARKs.-Status indeterminable. Possibly not a Notropis, but if so, most likely Notropis
chrysocephalus chrysocephalus. Original description date 31 December 1845.
=a questionable junior synonym of Notropis chrysocephalus chrysocephalus (Rafinesque)
Notropis ochoterenai Hubbs and Gordon 1937
See page 22.
Notropis xaenocephalus octoradius Baughman 1950
See page 22.
Alburus oligaspis Cope 1865
Cope 1865a:282-283. TYPE LOCALITY: "Kansas" (according to original description). This
was subsequently emended to "Platte River, near Fort Riley" (Cope 1865b). See REMARKS. LEC-
TOTYPE: ANSP 2753 (40.5), Dr. A. W. Hammond. Fowler (1910: pl. 20, fig. 48) designated lec-
totype by illustrating "type" (ANSP 2753); calculated length 40.0 mm SL. LECTOPARATYPE:
ANSP 2754 (1), paratopotype.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
REMARKs.-The type locality for this species is confused. Inasmuch as the Platte River does
not enter Kansas, the emended type locality given by Cope (1865b) cannot possibly be correct.
Cross (1967:226) wrote as follows about this: "Hammond was stationed temporarily at Fort Riley.
The fishes that he collected were delivered from that base to the Academy of Natural Sciences
of Philadelphia, but the precise localities of their capture are uncertain. Many were taken on an
expedition in 1856, through areas now within the states of Nebraska and Wyoming." Jordan
(1885a:122) was first to synonymize this form with Alburnus dilectus, which in turn was svnon-
ymized with N. atherinoides by Hubbs and Ortenburger (1929b:83). Original description date 13
February 1865.
= Notropis atherinoidcs Rafinesque
Notropis orcl Woolman 1894
\oolnan 1894:56. TYPE LO ALITY: Rio Grande, at El Paso, Texas. SYNTYPEs: CAS-SU 2278
(4, 51.7-61.7), USNM 212260 (ex CAS-SU 2278) (2, 56.9-63.4), A. J. Woolman, summer 1891.
REMARKS.-Condition of types very good to excellent. Counts: anal rays 8 in all specimens;
pharyngeal teeth 2,4-4,2 in one specimen examined. Miller (1976:11) noted differences between
individuals in the type series of .. simus and N. orca but indicated that these characters grade
together throughout the overall range. A possible subspecific relationship is also refuted by the
close proximity of their type localities (the Rio Grande in Texas and New Mexico, respectively).
Clark Hubbs (1957:6) recognized only N. simus from Texas, thereby tacitly synonymizing the
two. Original description date 2 May 1894.
= Notropi. simIus (Cope)
Codonma ornata Girard 1857
Girard 1857:195. TYPE LO( ALITY: Rio Chihuahua and tributaries, near Chihuahua City,
Chihuahua, Mexico. SYNTYPES: USNM 91 (2, 51.2-54.2), MCZ 1791 (ex USNM 91) (1, 50.6) John
Potts, 1855.
REMARKs.-Girard (1859a:53; pl. 29, figs. 22-25) listed four original syntypes and illustrated
one of these. Condition of syntypes soft, hut not bad for specimens of their age. Counts: teeth
4-4 (MCZ and one of USNM types); anal rays 7 (MCZ type). Teeth of USNM type in osteological
collection. Contreras (1975) and Miller (1976) tentatively placed this species in a separate genus,
Codota. Original description date 25 April 1857.
= Notropis ornatus (Cirard)
Notropis ortenburgeri Hubbs 1927
Hubbs, in Ortenburger and Hubbs 1927:127. TYPE LOCALITY: Mountain Fork R., 10 mi SE of
Broken Bow, McCurtain County, Oklahoma. HOLOTYPE: UMMZ 73092 (43.0), Univ. of Oklahoma
Mus. of Zool. field crew, June-July 1925.
REMARKS.-Original description date 1 January 1927.
= Notropis ortenburgeri Hubbs
Opsopoeodus osculus Evermann 1892
Evermann 1892:82-83. TYPE LOCALITY: Buffalo Bayou, 6 mi N of Houston, Harris County,
Texas. LECTOTYPE: USNM 45560 (43.0), B. W. Evermann, J. T. Scovell, and R. R. Gurley, 16
November 1891. Evermann and Kendall (1894: pl. 19, lowermost fig.) designated lectotype by
illustrating "type." LECTOPARATYPES: USNM 205205(3), paratopotypes; USNM 125143(36), CAS
13514 (old IU 9042) (4), CAS-SU 1127 (1). Evermann (1892:82) listed 82 original syntypes from
six localities, of which 40 cannot now be located.
REMARKS.-Gilbert and Bailey (1972:21-22) discussed status of this form. Original description
date 25 May 1892.
= Notropis emiliae emiliae (Hay)
Notropis oxyrhynchuts Hubbs and Bonham 1951
Hubbs and Bonham 1951:93-97; pl. 2 figs. 2-3. TYPE LOCALITY: Brazos R., Wellborn Cross-
BULLETIN FLORIDA STATE MUSEUM
ing, Brazos County, Texas. HOLOTYPE: UMMZ 129829 (50.8), Kelshaw Bonham and party, 21
October 1938. PARATYPES: UMMZ 129830 (230 originally, now 220), UF 14613 (ex UMMZ
129830) (10), paratopotypes; UMMZ 132054 (10), UMMZ 138238 (11), UMMZ 159363 (3), UMMZ
159364 (144), UMMZ 159365 (45), UMMZ 159366 (138), UMMZ 159367 (84).
REMARKS.-Original description date 30 March 1951.
= Notropis oxyrhynchus Hubbs and Bonham
Notropis ozarcanus Meek 1891
Meek 1891:129. TYPE LOCALITY: North Fork of White R., S of Cabool, Baxter County, Ar-
kansas. SYNTYPES: USNM 43230 (old U. S. Fish Comm. 3710) (4, 37.5-45.0), CAS 29305 (old
IU 8531) (7, 36.8-37.1), S. E. Meek, Louis Rettger, and F. M. Drew, 3 August 1889.
REMARKs.-A discrepancy arises regarding the number of syntypes of this species. Although
in his original description Meek (1891) listed only 10 specimens, the two type series combined
contain 11. The data on the original labels in the jars are in accord with Meek's description, but
the uniform condition of the types now present suggests they were all preserved at the same
time. Meek probably miscounted the number of specimens in preparing his original description.
Original description date 19 June 1891.
= Notropis ozarcanus Meek
Notropis emiliae peninsularis Gilbert and Bailey 1972
Gilbert and Bailey 1972:23-26; pls. Ib, 2b. TYPE LOCALITY: St. Johns R., at Fort Gates Ferry,
T12S, R26E, Ocala National Forest, Putnam County, Florida. HOLOTYPE: UMMZ 197672 (44.5),
C. L. Hubbs and party, 5 April 1941. PARATYPES: UMMZ 163356 (9), paratopotypes; UMMZ
106421 (5), UMMZ 106424 (1), UMMZ 101682 (2), UMMZ 126288 (3), UMMZ 158564 (36),
UMMZ 158637 (1), UMMZ 180659 (35), UF 179 (1), UF 180 (19), UF 228 (2), UF 546 (10), UF
4472 (1), UF 6258 (33), UF 6259 (12), UF 6260 (27), UF 6261 (35), UF 6262 (9), UF 6264 (42),
UF 6265 (42), UF 6266 (46), UF 6267 (52), UF 6268 (5), UF 6269 (35), UF 6270 (5), UF 6271 (13),
UF 6273 (2), UF 6274 (8), UF 6275 (146), UF 6276 (81), UF 6277 (11), UF 6279 (2), UF 6281 (4),
UF 6282 (2), UF 6284 (17), UF 6286 (67), FSU 1421 (3), FSU 1861 (2) (erroneously listed in orig-
inal description as 1661), FSU 2139 (3).
REMARKS.-Counts for holotype: pharyngeal teeth 5-4; anal rays 8; lateral-line scales 38;
caudal-peduncle circumferential scales 5-2-5= 12; predorsal scales 18; body circumferential
scales 12-2-11 = 26; pectoral rays 15-15; pelvic rays 8-8. Original description date 9 March 1972.
= Notropis emiliae peninsularis Gilbert and Bailey
Alburnellus percohromus Cope 1871
Cope 1871:440. TYPE LOCALITY: "St. Joseph, Missouri." (This locality probably is in eastern
Kansas, but may be central Missouri, as discussed below.) LECTOTYPE: ANSP 2993 (39.5), Dr.
William Stimpson. Fowler (1910: pl. 21, fig. 50) designated lectotype by illustrating "type"
(ANSP 2993); calculated length 40.1 mm SL. LECTOPARATYPES: ANSP 2994-3009 (13), parato-
potypes.
REMARKS.-The nomenclature and taxonomic status of this form have been confused. Jordan
(1885b:815) originally synonymized it with N. rubrifrons (= N. rubellus), which stood for 60 years
until Hubbs (1945:16-17) determined that N. percobromus was a valid species, most closely re-
lated to N. atherinoides. Hubbs and Bonham (1951:93) discussed the situation further, pointing
out that Hubbs and Ortenburger's (1929b:83-85) earlier acceptance of percobromus as the name
for the southwestern form of N. rubellus was in error, and resulted largely from Fowler's
(1910:290; pl. 21, fig. 50) description of N. rubrifrons, which was based in part on Cope's syn-
types of Alburnellus percobromus. Bailey and Allum (1962:56-60) disputed Hubbs' (1945) recog-
nition of N. percobromus as a valid species, and synonymized it with N. atherinoides.
I reexamined the 14 syntypes of A. percobromus which, although slightly soft, are in good
condition for old specimens. All are N. rubellus. Especially diagnostic are the small eye, sharply
pointed snout, relative positions of the dorsal and pelvic fins (dorsal fin more posteriorly situ-
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
ated in N. rubellus), number of anal-fin rays (10 in 11 of the 14 types vs. usually 11 in atheri-
noides), and number of body-circumference scales (26 to 28), which are higher than the counts
(23 to 25) usually found in N. atherinoides (Snelson 1968:792). Thus Jordan's (1885b) original
synonymization of percobromus with rubrifrons was correct, as was Hubbs and Ortenburger's
(1929b) application of the name percobromus to the southwestern population of rubellus. Al-
though N. percohromus (as now understood) may well be a valid form, either as a species or sub-
species, final judgment must await further study. It is tentatively regarded here as a junior syn-
onym of N. rubellus.
The type locality for Alburnellus percobromus probably is in error, as St. Joseph, Missouri,
is outside the present range of N. rubellus (Cross 1967:104; Pflieger 1971:485, map 46). On the
first page of the paper in which percobromus was described, Cope (1871:432) stated "a few
species from the neighborhood of St. Joseph, Northwestern Missouri, are added." This indicates
that the locality "St. Joseph, Missouri," which subsequently appeared in the account of per-
cobromus, should not be taken literally. N. rubellus occurs today in Kansas within about 75 miles
of St. Joseph; the closest locality in Missouri is slightly over 100 miles away. Inasmuch as Kansas
localities for this species are slightly closer, it is more likely that the original specimens came from
that state (if so, presumably from the Kansas River drainage) rather than from Missouri. Possibly
a detailed morphological comparison of populations from these areas with the types will help
resolve this problem. Original description date (31 December) 1871.
= Notropis rubellus (Agassiz)
Notropis perpallidus Hubbs and Black 1940
Hubbs and Black 1940:46-49. TYPE LOCALITY: Saline R., 5 mi N of Warren, Bradley County,
Arkansas. HOLOTYPE: UMMZ 125991 (30.0), John D. and Ruby Y. Black, 20 June 1939. PARATYPE:
UMMZ 125992 (1), paratopotype.
REMARKS.-Snelson and Jenkins (1973) reviewed this species. They showed (1973:293) that the
name N. ionthas, which was earlier introduced by Jordan and Gilbert (1886:11), was based on
this species, but as it appeared as a junior synonym and was not used subsequently, it has no legal
nomenclatural standing (International Code, articles 1 (d) and 16(b)(ii)). Original description date
30 March 1940.
= Notropis perpallidus Hubbs and Black
Notropis petersoni Fowler 1942
Fowler 1942:1-2, figs. 1-2. TYPE LOCALITY: Crane Cr., below U.S. hwy. 1 bridge, Moore Co.,
North Carolina. HOLOTYPE: ANSP 69967 (50.0), C. Bernard Peterson, G. A. Coventry, and F. A.
Ulmer, Jr., 23 August 1940. PARATYPEs: ANSP 69968 (1), paratopotype.
REMARKS.-Swift (1970) reviewed this species. Original description date 16 September 1942.
= Notropis petersoni Fowler
Hybopsis phaenna Cope 1865
Cope 1865a:279. TYPE LOCALITY: tribes. of Delaware R., near Trenton, New Jersey. LECTO-
TYPE: ANSP 4389 (85.0), Dr. Charles C. Abbott. Fowler (1910: pl. 16, fig. 15) designated lecto-
type by illustrating "type" (ANSP 4389); calculated length 86.8 mm SL. LECTOPARATYPES: ANSP
4390-4391 (2), paratopotypes.
REMARKS.-Jordan (1876b:281) was first to synonymize this form with Hybopsis (= Notropis)
hudsonius. Original description date 13 February 1865.
= Notropis hudsonius (Clinton)
Notropis phenacobius Forbes 1885
Forbes 1885:137. TYPE LOCALITY: Illinois R., at Peoria, Illinois. SYNTYPES: None located. Ten
specimens were originally present in the INHS collection, but nine of these were lost prior to
1920 (Forbes and Richardson 1920:139). The remaining type was not listed by Smith and Bridges
(1960), and now is also presumed to be lost. The absence of any types of this species from the
1978
BULLETIN FLORIDA STATE MUSEUM
INHS collection was recently reconfirmed by P. W. Smith (in litt., 4 February 1974). Six speci-
mens catalogued as N. phenacobius and labelled as questionable types are in the Field Museum
of Natural History (FMNH 70579). The provenance of these specimens cannot be determined,
and their identification (by me) as N. stramineus is at variance with the original description of
N. phenacobius (see REMARKS). I conclude, therefore, that the six specimens in question are not
part of the original syntypic series.
REMARKS.-Jordan and Evermann (1896:263) were first to synonymize this form with N. scylla
(=N. stramineus), an action that has since been followed by others (e.g. Suttkus 1958:313). Re-
examination of the original description of N. phenacobius, as well as the subsequent description
and illustration (head only) by Forbes and Richardson (1920:138-139, fig. 33), strongly suggests
that this form is, in fact, a senior synonym of Notropis amnis Hubbs and Greene 1951.
First, the descriptions in Forbes (1885) and Forbes and Richardson (1920) agree well with
that of N. amnis in nearly all important features (mouth small, inferior, and horizontal; nose
long; head square; eye unusually large; anal rays I-8; dorsal fin before ventrals; lateral-line
scales 35 to 36; predorsal scales 13 to 14). These characters could also apply to N. stramineus,
with the one important exception of anal-ray count (almost always 7 in that species). The illus-
tration of the head in Forbes and Richardson (1920:138, fig. 33) bears a much stronger re-
semblance to N. amnis than to N. stramineus, with the tiny, horizontal, inferior mouth being
particularly diagnostic. The one morphological character at variance with that usually found in
N. amnis is pharyngeal-tooth count, which was recorded by Forbes (1885) and Forbes and
Richardson (1920) as 4-4. Although the usual pharyngeal-tooth count in N. amnis is 1,4-4,1, the
tooth in the lesser row is small and fragile (thus easily broken), and sometimes is missing alto-
gether (Glenn H. Clemmer, pers. comm.); this could easily account for the apparent discrepancy
in count.
Second, Forbes and Richardson were familiar with N. stramineus in Illinois, inasmuch as they
treated that species (under the name N. blennius) in the account immediately preceding that of
N. phenacobius. It might be argued that N. phenacobius is a synonym of N. colucellus, a species
usually having eight anal rays and which otherwise shares certain features with N. amnis. As
N. volucellus is common and widespread throughout the midwestern United States and was not
separated specifically from the very similar appearing N. stramineus until later (Hubbs and
Greene 1928:375-379), it is far more likely that Forbes and Richardson's "N. blennius" actually
was a complex of N. stramineus and N. volucellus.
Finally, the rarity in Illinois of N. phenacobius, which was known to Forbes and Richardson
only from the 10 syntypes from the Illinois River at Peoria, is concordant with the rarity of N.
amnis from the area. Hubbs and Greene (in Hubbs 1951a:7) recorded amnis only from two
widely separated localities in the Illinois River drainage: the Sangamon River in Champaign
County, Illinois; and the Iroquois River (above Peoria) in northwestern Indiana. The type lo-
cality for N. phenacobius thus is within the known geographic range of N. amnis.
In summary, circumstantial evidence strongly indicates that N. phenacohius Forbes 1885 is
not a junior synonym of N. stramineus (Cope 1865), as previously thought, but rather is a senior
synonym of N. amnis Hubbs and Greene 1951. Three courses of action are open under these
circumstances: (1) change the specific name of amnis to phenacobius, (2) petition the Interna-
tional Commission to rule the name phenacobius unavailable in the genus Notropis, or (3) post-
pone action of any kind, as no types of N. phenacobius are known to exist and thus its identity
cannot be categorically established. The last course of action is the one presently being followed.
Original description date (31) March 1885.
= a probable senior synonym of Notropis amnis Hubbs and Greene
Squalius photogenis Cope 1865
Cope 1865a:280. TYPE LOCALITY: Youghiogheny R., Pennsylvania. LECTOTYPE: ANSP 22280
(39.0), E. D. Cope (specimen lost). Fowler (1909:544-545) designated lectotype and subsequently
illustrated the "type" (Fowler 1910: pl. 21, fig. 51). Standard length of lectotype calculated
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
from Fowler's drawing. See Gilbert (1971a) for further remarks on lectotype designation. LECTO-
PARATYPE: ANSP 22281 (1). This specimen cannot presently be located. For further discussion
see REMARKS.
REMARKS.-This species has had an extremely complicated nomenclatural history, which I
reviewed earlier (Gilbert 1971a:477-478). Thus only a brief summary is necessary here.
Despite the indication above, both specimens in the type jar (ANSP 22280-22281) are N.
spilopterus. I (1971a) assumed this to have resulted from misplacement of the syntypes, inas-
much as Fowler's (1910: pl. 21, fig. 51) illustration of ANSP 22280 is clearly not N. spilopterus,
but rather is a reasonably good representation of N. photogenis as presently recognized. The true
syntypes (actually the lectotype and lectoparatype) cannot be located, but may still be present
in the ANSP collection. Finally, reexamination of Cope's (1865a:280) original account definitely
shows that N. photogenis, and not N. spilopterus, was the species on which his description was
based. Original description date 13 February 1865.
= Notropis photogenis (Cope)
Notropis pilsbryi Fowler 1904
Fowler 1904b:245-246, fig. TYPE LOCALITY: White R. system, at Rogers, Benton County,
Arkansas. HOLOTYPE: ANSP 24514 (72.5), H. A. Pilsbry, April 1903. PARATYPES: ANSP 24515-
24520 (6), paratopotypes.
REMARKS.-This species was synonymized with the closely related N. zonatus by Hubbs and
Ortenburger (1929b:81-82), as well as by Hubbs and Brown (1929:37). Hubbs and Moore (1940)
subsequently elevated it to a subspecies of N. zonatus, where it remained until I (Gilbert
1964:133-136) restored it to full species status. The fish listed by Forbes and Richardson (1920:149)
as N. pilshryi was based on a hybrid: N. cornutus x N. rubellus (Hubbs and Brown 1929:37;
Hubbs and Moore 1940:96). Original description date 7 April 1904.
= Notropis pilsbryi Fowler
Notropis amnis pinnosa Hubbs and Bonham 1951
Hubbs and Bonham, in Hubbs 1951a:17-25; pl. 1, figs. 3-5. TYPE LOCALITY: Trinity R., Mag-
nolia Point, 10 mi SW of Palestine, Anderson County, Texas. HOLOTYPE: USNM 45556 (45.0),
B. W. Evermann, R. R. Gutley, and J. A. Singley, 23 November 1891. This specimen is also the lec-
totype of N. nocomis Evermann 1892. PARATYPEs: USNM 93517 (20), paratopotypes; USNM
36475 (1), USNM 69335 (1), USNM 93514 (7), UMMZ 160277 (7). The series USNM 36475 and
69335 were not listed as paratypes by Hubbs (1951a), but are so indicated in the USNM fish
catalogue. Other paratypes were said to be present in the University of Texas and Texas A & M
collections.
REMARKS.-This form was previously described by Evermann (1892:78-79) as N. nocomis.
However, the specific name nocomis had earlier been used (Jordan and Gilbert 1886:23-24) for
a southwestern population of N. colucellus, although it first appeared as a nomen nudum and
thus had no legal status. Hubbs' (1951a:11-12) subsequent action provided such status, with the
result that N. nocomis Evermann 1892 was now preoccupied and required a new name, which
Hubbs and Bonham proposed. Recent studies by Clemmer (1970) indicate the subspecies pinnosa
is not valid. Original description date 14 March 1951.
= Notropis amnis Hubbs and Greene
Photogenis piptolepis Cope 1871
Cope 1871:438-439. TYPE LOCALITY: Red Cloud Cr., tributary to North Platte R., (Wyo-
ming?). TYPES: None located.
REMARKS.-Although the types of Photogenis piptolepis are apparently lost, study of the orig-
inal description, together with consideration of the type locality, clearly indicates this is the
species presently called N. dorsalis. The form piptolepis may be a valid subspecies, but final
clarification must await further study. Original description date (31 December) 1871.
= Notropis dorsalis (Agassiz)
1978
BULLETIN FLORIDA STATE MUSEUM
Rutilus plargyrus Rafinesque 1820
Rafinesque 1820b:50-51. TYPE LOCALITY: "Kentucky." TYPES: None located.
REMARKs.-The identification of this form cannot be determined with certainty. The species
to which the rather ambiguous description most logically applies is N. chrysocephalus, and I re-
ferred it questionably to that species (Gilbert 1964:158). Original description date (31) May 1820.
= possibly Notropis chrysocephalus chrysocephalus (Rafinesque)
Alburnops plumbeolus Cope 1865
Cope 1865a:282. TYPE LOCALITY: Saginaw R., Flint, Michigan. LECTOTYPE: ANSP 2061
(40.5), E. D. Cope. Gilbert (1964:160) designated lectotype (as ANSP 2055). For additional dis-
cussion see REMARKS below. LECTOPARATYPES: ANSP 2056-2060 (5), paratopotypes. The seventh
original syntype (ANSP 2055) is N. heterodon, and has been recatalogued under that name (also
see discussion below).
REMARKs.-Although I (Gilbert 1964:160) designated the lectotype indicated above (as ANSP
2055), it was subsequently found that Fowler (1910: pl. 18, fig. 30) had made an earlier lectotype
designation by illustrating the "type" (ANSP 2055). Unfortunately, his illustrated specimen, the
largest in the type series, proves to be the lone individual of N. heterodon (Cope), a species that
was originally described as Alburnops heterodon in the same paper as A. plumbeolus. Should
Fowler's lectotype designation be allowed to stand, plumbeolus and heterodon would become
synonyms, and, as they were described simultaneously, the former name conceivably could sup-
plant the latter, depending upon the action of the first reviser. As Cope's original description of
A. plumbeolus clearly was based on N. chrysocephalus (represented by the other six specimens
in the syntypic series) and not N. heterodon, I (Gilbert 1977) have petitioned the International
Commission to suppress Fowler's lectotype designation. As both Fowler and I used the same
catalogue number (ANSP 2055) for two different specimens, I have also requested that this num-
ber henceforth be restricted to Fowler's lectotype, and that mine be changed to ANSP 2061.
Original description date 13 February 1865.
= Notropis chrysocephalus chrysocephalus (Rafinesque)
Notropis potteri Hubbs and Bonham 1951
Hubbs and Bonham 1951:100-109; pl. 3, figs. 1-3. TYPE LOcALITY: Waco Cr., near Waco,
McLennan County, Texas. HOLOTYPE: UMMZ 120228 (45.3), C. L. Hubbs and family, Leo T.
Murray, 21 June 1938. PARATYPES: UMMZ 120227 (12 originally, now 10; 22 erroneously listed
in original description), USNM 117568 (ex UMMZ 120227) (2), paratopotypes; UMMZ 92109
(1), UMMZ 92131 (1), UMMZ 101801 (1), UMMZ 110531 (2), UMMZ 110592 (3), UMMZ 110593
(1), UMMZ 112477 (2), UMMZ 129831 (68), UMMZ 129843 (1), UMMZ 132051 (18), UMMZ
159374 (102), UMMZ 159375 (24, plus 10 to G. E. Potter), UMMZ 159376 (34).
REMARKS.-This species first appeared (under the above name), as a labelled illustration only,
in Potter's (1938) zoology textbook, and by name only in Baughman's (1950) checklist of Texas
fishes. Neither qualifies as a valid description, according to Article 25, Item C, of the Interna-
tional Rules. Suttkus and Clemmer (1968:35-38) discussed the status and relationships of this
species. Original description date 30 March 1951.
= Notropis potter Hubbs and Bonham
Hybognathus procne Cope 1865
Cope 1865a:283. TYPE LOCALITY: Conestoga R. tribe to Susquehanna R.), Pennsylvania.
LECTOTYPE: ANSP 3152 (41.0), Jacob Stauffer. Fowler (1909:532) designated lectotype, although
it should be noted that in the text and illustration in his later paper he (Fowler 1910:275; pl. 15,
fig. 8) referred only to "cotypes." LECTOPARATYPES: ANSP 3153-3162 (9), paratopotypes.
REMARKS.-Condition of types poor, but pertinent characters evident. Counts: pharyngeal
teeth 4-4 and anal rays 7 in all specimens examined. Fowler (1910:275) listed 11 syntypes, but
only 10 of these now seem to be extant. He also illustrated a "cotype" (Fowler 1910: pl. 15, fig.
8); estimated standard length 40.5 mm. This is very close to the largest individual present in the
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
jar of syntypes, and this specimen (which I have segregated from the remainder of the series)
is considered to be the lectotype. Original description date 13 February 1865.
= Notropis prone (Cope)
Moniana proscrpina Girard 1857
Girard 1857:2(X). TYPE LOCALITY: Devils R., probably just above mouth into Rio Grande,
Val Verde County, Texas. SYNTYPES: USNM 117 (9[10 listed in USNM catalogue], 29.8-45.8, plus
one right pharyngeal arch in osteological collection). ANSP 3026 (ex USNM 117) (1, 39.0), MCZ
1688 (ex USNM 117)(2, 33.7-37.7), BMNH 1883.12.14.216(ex USNM 117)(1, 40.1), John H. Clark,
1851.
REMARKs.--irard (1859a:59) listed 10 syntypes for USNM 117, but the total number of ex-
tant syntypes is 12. Moniana proserpina and Moniana aurata, which were described in the same
paper and on the same page by Cirard (1857), were first synonymized (under the former name)
by Jordan i I i !'I., who thus qualifies as first reviser. This action was later confirmed by
Huhbs and Ortenburger (1929b:75), following examination of types. Condition of types fairly
good for old specimens. Counts: pharyngeal teeth 4-4, anal rays 8 in both ANSP and MCZ svn-
types. Lytle (1972) reviewed species. Original description date 25 April 1857.
= Notropis proserpinus (Girard)
Moniana pulchella Girard 1857
Cirard 1857:2(X). TYPE L( ALITY: Sugar Loaf Cr., tributary to Poteau R., about 20-25 mi S
Fort Smith, Arkansas, vicinity of Poteau, Oklahoma. Locality is very near the Oklahoma-
Arkansas state line and could be either in Le Flore County, Oklahoma, or Sebastian County,
Arkansas. SYNTYPES: None located. Girard (1858:276) listed a single specimen (UlSNM 122) from
the above locality, collected by H. B. Mollhausen in 1853, but this apparently is no longer ex-
tant.
REMARKs.-Some confusion attends the status of the type specimens of Moniana pulchella.
Cirard (1858:276) listed, in addition to the single specimen (USNM 122) above, a series of 18
specimens (USNM 121), collected by Dr. George G. Shumard in 1853 from the Arkansas River
near Fort Smith, Arkansas. Although not specifically stated in Cirard's 1858 paper, these speci-
mens conceivably could be presumed to comprise a second syntypic series. They have been so
labelled in the jars (the original series was divided between the USNM, ANSP, MCZ, and pos-
sibly other collections) and were also regarded as types byv Hubbs and Ortenburger (1929b:74),
who studied the two individuals now in the MCZ. It is possible that Girard did, in fact, intend
USNM 121 to represent a second type series; unfortunately, they cannot be so regarded, inas-
much as neither they nor the locality from which they came were mentioned in the original de-
scription of the species (Girard, 1857:2(X)). Girard (1858: pl. 58, figs. 11-15) illustrated M. pul-
chella, but there is no indication from which series the specimen came on which the illustration
is based.
The specimens still remaining from USNM 121 are recorded here, although they are no
longer regarded as valid syntypes: USNM 121 (three pharyngeal arches only, in osteological
collection), ANSP 4085 (ex USNM 121) (1, 45.5), MCZ 1797 (ex USNM 121) (2, 43.5-48.2). Counts:
pharyngeal arches 4-4 (1), 4-? (USNM 121); 4-4 (ANSP 4085); 4-4 and 0,44,1 (MCZ 1797); anal
rays 9 in all three specimens examined.
Jordan (1885b:812) was first to synonymize this form with AV. lutrensis, an action Hubbs and
Ortenburger (1929b:74) confirmed after examining the presumed syntypes in the MCZ. Original
description date 25 April 1857.
= Notropis lutrensis (Baird and Girard)
Minnilus punctulat.us Hay 1881
Hay 1881:508-509. TYPE LOCALITY: tributary to Tuscuimbia R., near Corinth, Alcorn County,
Mississippi. LECTOTYPE: USNM 27430 (38.6), 0. P. Hay, March-April 1880. Snelson and Pflieger
(1975:235) designated lectotype. LECTOPARATYPES: C(AS-SU 4076 (4), paratopotypes.
REMARKS.-This form was regarded as a questionably valid subspecies (Jordan and Ever-
BULLETIN FLORIDA STATE MUSEUM
Vol. 23, No. 1
mann 1896:301-302) or species (Jordan, Evermann, and Clark 1930:126) for many years. It ap-
parently was first formally synonymized with N. ionhratilis by Cook (1959:122). Original descrip-
tion date 23 February 1881.
= Notropis imnbrtilis cyatnocephaIlus (Copeland)
Photogenis pyrrhoimelas Cope 1870
Cope 1870:463-464. TYPE LOCALITY: tributary of upper Catawba R., North Carolina. LE:-
TOTYPE: ANSP 2631 (66.0), E. D. Cope, Autumn 1869. Fowler (1910: pl. 18, fig. 29) designated
lectotype by illustrating "type" (ANSP 2631); calculated length 69.2 mm SL. LECTOPARATYPES:
ANSP 2632-2725 (94), paratopotypes. Counts for lectotype: pharyngeal-teeth 1,4-4,1: anal rays
10. Gibbs (1955) reviewed species. Original description date 21 November 1870.
= Notropis pyrrhomelas (Cope)
Notropis rasconis Jordan and Snyder 1899
See page 22.
Notropis heterolepis regalis Hubbs and Lagler 1949
Hubbs and Lagler 1949:119-123; pl. 2, fig. 3. TYPE LOCALITY: Harvey Lake, Isle Royale,
Keweenaw County, Michigan. HOLOTYPE: UMMZ 100027 (81.0), Walter Koelz, 1 September
1929. PARATYPES: UMMZ 1(XK)28 (108), paratopotypes; UMMZ 144474 (23).
REMARKs.-The subspecies regalis is not generally recognized, although it has never been
formally synonymized. Original description date (28) February 1949.
= Notropis heterolepis Eigenmann and Eigenmann
Notropis reticulatus Eigenmann and Eigenmann 1893
Eigenmann and Eigenmann 1893a:152-153. TYPE LO(ALITY: Fort QO'Appelle, Saskatchewan;
and Assiniboine R., Brandon, Manitoba. SYNTYPES: USNM 64727 (1,33.5), BMNH 1892.12.30.533-
542 (10, 36.0-46.0), Brandon; USNM 125158 (1, 38.6), BMNH 1892.12.30.560-567 (8, 35.0-53.5),
CAS-SU 541 (1, 40.6), Fort Qu'Appelle, C. H. Eigenmann, 1892.
REMARKS.-A total of 13 and 24 specimens were originally recorded from Brandon and Fort
Qu'Appelle, respectively. Counts for USNM 64727 and USNM 125158: pharyngeal teeth 4-4 in
both; anal rays 7 in both. Hubbs and Greene (1928:375-376) were first to synonymize N. reticu-
latus with N. deliciosius (= N. stramineus). Original description date 4 February 1893.
= Notropis stramineus (Cope)
Notropis heterodon richardsoni Hubbs and Greene 1926
Hubbs and Greene, in Hubbs 1926:39-40; pl. 3, fig. 2. TYPE LOCALITY: Fox R., opposite Lock
25, Columbia County, Wisconsin. HOLOTYPE: UMMZ 66536 (44.0), C. Willard Greene and
Harmon S. Jones, 26 August 1925. PARATYPES: UMMZ 66537 (34) (seven additional specimens,
originally included among paratypes, were subsequently reidentified as N. chalyhaeus and re-
moved by C. C. Swift), USNM 93426 (ex UMMZ 66537) (1), paratopotypes; UMMZ 73436 (1),
UMMZ 73450 (3), UMMZ 73493 (16), UMMZ 73524 (2), UMMZ 73574 (1), UMMZ 73632 (1),
UMMZ 73637 (7), UMMZ 73662 (12), UMMZ 73684 (2), UMMZ 73701 (31), UMMZ 73712 (6),
UMMZ 73730 (1), UMMZ 73800 (2), UMMZ 73826 (1), UMMZ 73857 (2), UMMZ 73912 (9).
REMARKs.- Bailey, Winn, and Smith (1954:126) were first to synonymize this form with N.
roseurs (= N. texanns). Original description date 7 July 1926.
= Notropis texan us (Girard)
Notropis robhstus Meek 1902
Meek 1902:82-83; pl. 19, upper fig. TYPE LO:ALITY: Rio Conchos, Jimenez, Chihuahua, Mex-
ico. HOLOTYPE: FMNH 3548 (46.8), S. E. Meek and F. E. Lutz, April-June 1901. PARATYPES:
FMNH 3549 (16), paratopotypes; FMNH 3534 (3).
REMARKS.- Hubbs and Hubbs (1958:3(X)) were first to synonymize this form with N. bray-
toni. Original description date (31) May 1902.
= Notropis braytoni Jordan and Evermann
GILBERT: NOTROPIS TYPE CATALOGUE
Notropis roscipinnis Hay 1885
Hay, in Jordan 1885b:815. TYPE LOCALITY: Chickasawha R. and tributaries), at Enterprise,
Clarke County, Mississippi. LECTOTYPE: USNM 27420 (43.0), O. P. Hay, March-April 1880. Snel-
son (1972:46) designated lectotype. LECTOPARATYPES: USNM 203325 (16), paratopotypes. One
additional specimen from the original syntypic series represents an unidentified species of .o-
tropis (not N. roseipinnis) and is not included among the lectoparatypes.
REMARKS.-The species name roseipinnis is a replacement name for Minnilus ruhripinnis Hav
1881, which is preoccupied in the genus Notropis. According to Article 72d, International Code,
the types for rubripinnis must also serve as the types for the replacement name. Snelson (1972:45-
58) reviewed species. Original description date 2 October 1885.
= Notropis roseipinnis Hay
Luxilus roscus Jordan 1877
Jordan 1877e:61. TYPE LOCALITY: Natalbanv R., near Tickfaw, Tangipahoa Parish, Louisi-
ana. LECTOTYPE: USNM 17831 (53.5), T. H. Bean and O. P. Maxson, December 1876. Suttkus
and Raney (1955c:31) designated lectotype. LECTOPAHATYPES: USNM 171788 (7). Three other
species, comprising a total of 11 specimens, were included in original svntvpic series. Two other
specimens (MCZ 35908) from the original svntypic series have also been examined: neither is
N. roses (= N. texanus) as presently understood.
REMARKs.-Sunttkus (1958) reviewed the status of Luxilurs roses in relation to its senior svn-
onym Cyprinella texana Girard 1857. Original description date 131 December) 1877.
= A.otropis texanus (Girard)
Alburnus rnuhellus Agassiz 1850
Agassiz 1850:364-366; pl. 3, figs. 1-3. TYPE LOCALITY: Lake Superior, at Sault Ste. Marie,
either Michigan or Ontario. HOLOTYPE: MCZ 1749 i58.0), L. Agassiz, 27 June 1848. PARATYPES:
MCZ 1749A (4), UMMZ 871() (ex MCZ 1749A) (1), paratopotypes.
REMARKs.-AlIbirn.us rubellus was long regarded a junior synonvm of X. athcrinoides Rafi-
nesque 1818 (see Jordan 1878a:422). Hubbs and Brown (1929:35), in clarifying its status, pointed
out that the nine specimens listed by Agassiz (1850) from "the Pic" are N. atherinoides. whereas
the five from Sault Ste. Marie are N. rubellus. One specimen from the latter locality, which was
tintagged and separated from the other types, clearly formed the basis (as determined by a physi-
cal abnormality) for Agassiz s description and illustration. It was consequently regarded bv Hubbs
and Brown as the holotype of Alburmnus rubellus. The holotype is bleached and the body soft,
but otherwise is in relatively good condition for a specimen its age. Counts for holotvpe: pharyn-
geal teeth, 2,4-4,2; anal rays 10. Original description date (31 December) 1850.
= Notropis rubellus (Agassiz)
Rutilus? ruber Rafinesque 1820
Rafinesque 1820h:52. TYPE LOCALITY: tributaries of Elkhorn Cr. (Kentucky R. drainage),
near Lexington, Kentucky. TYPES: None located.
REMARKs.-This form cannot be identified with certainty, particularly as Rafinesque
(1820h:52), in the original description, stated that he personally had not yet seen the fish. Snel-
son and Pflieger (1975:233) suggested that it might be N. umbratilis, but the closely related N.
gardens is abundant in the drainage, and Jordan (1877d:32) believed this was the most likely pos-
sibility. Other possibilities are N. rubellns and Phoxinus erythrogaster. Original description date
(31) May 1820.
= Not definitely identifiable, but most likely Notropis artdns.
Hyhopsis rubricroceus Cope 1868
Cope 1868h:231-232; pl. 24, fig. 4. TYPE LO:ALITY: near mouth of Tumbling Cr., trib. to
North Fork of Holston R., Virginia. LECTOTYPE: ANSP 2907 (58.5), E. D. Cope, October 1867.
Fowler (1910: pl. 19, fig. 34) designated lectotype by illustrating "type" (ANSP 2907); calculated
length 59.3 mm SL. LECTOPARATYPES: ANSP 2908-2917 (10), paratopotvpes. Remaining three
1978
BULLETIN FLORIDA STATE MUSEUM
specimens in original syntypic series (N. chrysocephalus) have been removed and recatalogued
under that name.
REMABKs.-Condition of types fair, with all important features present. Counts: pharyngeal
teeth 2,4- (left arch only examined) in four specimens checked; anal rays 8 in 10, 9 in 1. Original
description date 2 December 1868.
= Notropis rubricroceus (Cope)
Alburnus rubrifrons Cope 1865
Cope 18651:85. TYPE LOCALITY: Kiskiminitas R. (trib. to Allegheny R.), Pennsylvania. LEC-
TOTYPE: ANSP 4035 (50.0), E. D. Cope. Fowler (1909:543) designated lectotype and later (Fowler
1910: pl. 21, fig. 49) also illustrated the "type" (ANSP 4035); calculated length 45.9 mm SL. LEC-
TOPARATYPES: ANSP 4036-4039 (4), MNHN 2682 (1), paratopotypes.
REMARKS.-Some confusion exists regarding Fowler's lectotype designation of Alburnus rubri-
frons. Although Fowler (1909:543) designated and subsequently illustrated the "type" (Fowler
1910: pl. 21, fig. 49), he made no such reference in the text of the latter paper (1910:290), in
which he merely referred to five "cotypes." The estimated length of the individual illustrated is
only 45.9 mm SL, which is shorter than at least three of the five syntypes (range 48-50 mm SL).
(The heads are broken off the other two.) The discrepancy between calculated and actual length
of the largest individual is more than for most specimens Fowler illustrated in his 1910 paper,
but it is not great enough to cast serious doubt that the illustration of A. rubrifrons (1910: pl. 21,
fig. 49) was based on one of the types. Counts for original syntypes: pharyngeal teeth 2,4- (one
specimen); anal rays 10 (three specimens). Hubbs and Brown (1929:34-35) were first to synony-
mize this species with N. rubellus. Original description date 7 August 1865.
= Notropis rubellus (Agassiz)
Nocomis rubrifrons Jordan 1877
See pages 21,92.
Argyreus rubripinnis Heckel 1843
Heckel 1843:1040, fig. pharyngeall teeth only). TYPE LOCALITY: "Boreal America." TYPES:
None located. Specimens said to have been sent to MNHN "par Msr. Milbert de New-York,"
but Jordan (1880a) did not list them as being there.
REMARKS.-Girard (1857:196) was first to synonymize this form with Plargyrus corntuss
(= N. cornutus). Original description date (31 December) 1843.
= Notropis cornutus (Mitchill)
Cyprinella rubripinna Carman 1881
Garman 1881:91. TYPE LOCALITY: Lago del Muerto, near Parras, Coahuila, Mexico. SYNTYPES:
MCZ 24891 (2, 53.0-56.0) MCZ 24892 (20, 27.8-54.0), UMMZ 86663 (ex MCZ 24892) (6, 40.0-
56.0), USNM 120257 (6, 35.5-51.3), Dr. E. Palmer, 1880.
REMARKS.-As the species name rubripinnis was preoccupied in Notropis by Argyrcus ruhri-
pinnis Heckel 1843, Jordan (18851:813) proposed garmani as a substitute name for Cyprinella
rubripinna. According to Article 72d of the International Code, the types for the last species
must also serve as the types for the replacement name. For comments on and counts for the syn-
types, see N. ganrani. Original description date 31 January 1881.
= Notropis lutrensis garmani Jordan
Minnilus rubripinnis Hay 1881
Hay 1881:509-510. TYPE LO(ALITY: Chickasawha R. (and trials) Enterprise, Clarke County,
Mississippi. LECTOTYPE: USNM 27420 (43.0), O. P. Hay, March-April 1880. Snelson (1972:46)
designated lectotype. LECTOPARATYPES: USNM 203325 (16), paratopotypes. One additional speci-
men from the original syntypic series represents an unidentified species of Notropis (not N. rosei-
pinnis). and is not included among the lectoparatypes.
REMARKS.-As the species name rubripinnis is preoccupied in Notropis, Hay (in Jordan
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
1885b:815) proposed roseipinnis as a substitute name. According to Article 72d of the Interna-
tional Code, the types for Minnilus rubripinnis must also serve as the types for the replacement
name. Original description date 23 February 1881.
= Notropis roseipinnis Hay
Moniana rutila Girard 1857
Girard 1857:201. TYPE LOCALITY: Rio Monterrev tribe to Rio San Juan), (adereita (= Ca-
dereyta) Nuevo Leon, Mexico. SYNTYPES: USNM 95 (two sets of pharyngeal arches in osteologi-
cal collection), MCZ 1794 (ex USNM 95) (1, 46.6), ANSP 4086 (ex USNM 95) (1, not measurable
[In several pieces]), Lt. D. N. Couch, 1853.
REMARKS.-Girard (1859a:58) listed 10 original syntvpes (USNM 95), but only those listed
above can now be found. Condition of MCZ 1794 relatively good for its age. Specimen with
heavy tuberculation on top of head, none before or below eves. Counts for MCZ 1794: pharyn-
geal teeth 4-4; anal rays 8. Counts for USNM 95: pharyngeal teeth 4-4. Lytle (1972), who reviewed
species, said N. rutilus is most closely related to N. proserpinus. Original description date 25
April 1857.
= Votropis rutilus (Girard)
Notropis sabinae Jordan and Gilbert 1886
Jordan and Gilbert 1886:15. TYPE LOCALITY: Sabine R., 5 mi S of Longview, Gregg County,
Texas. SYNTYPES: USNM 36484 (5, 19.8-39.8), CAS-SU 1071 (1, 34.3), D. S. Jordan, C. H. Gilbert,
Joseph Swain, and S. E. Meek, July-September 1884.
REMARKS.-Original description date 17 September 1886.
= Notropis sabinae Jordan and Gilbert
Notropis sahladonis Hubbs and Hubbs 1958
Hubbs and Hubbs 1958:297-307, fig. 1. TYPE LOCALITY: Rio Salado, 1 mi W of Manuel Mar-
tinez Parela and 10 mi S of Anahuac, Nuevo Leon, Mexico. HOLOTYPE: UMMZ 173132 (32.0),
Luciano Val Guerra, 11 June 1951. PARATYPES: UMMZ 173133 (7), paratopotypes; UMMZ
130349 (5), UMMZ 173134 (2), USNM 175276 (12), FMNH 62160 (11), FMNH 62169 (7), CAS-
SU 50173 (6), TNHC 1680 (16), TNHC 1722 (1), IPN (uncat.) (31).
REMARKS.-Original description date 22 December 1958.
= Notropis saladonis Hubbs and Hubbs
Ceratichthys sallaci Gunther 1868
Giinther 1868:484. TYPE L)OCALITY: Cuernavaca, Mexico. SYNTYPES: BMNH 1868.3.3.1-3
(3, 64.0-72.0), M. Boucard.
REMARKS.-Species named for Sefnor Sall&. Counts: pharyngeal teeth 4-4: anal rays 7. For
further remarks on this species see Miller (1976:10). Original description data 14 March 1868.
= Notropis sallei (Giinther)
Alburnops saludlanus Jordan and Brayton 1878
Jordan and Brayton 1878:16-18. TYPE LO ALITY: Saluda R., at Farr's Mill, west of Green-
ville, Greenville County, South Carolina. SYNTYPES: USNM 31128 (4, 66.0-87.5), D. S. Jordan, A.
W. Brayton, C. H. Gilbert et al., summer 1877.
REMARKS.-This form has frequently been regarded as a valid subspecies of N. hudsonius,
and differs from typical hudsonius in several ways pharyngeall tooth count, development of
caudal spot, head and body depth), but Seaman (1968) found that the perplexing morphological
variation in the region where the two forms come together makes even a subspecific differentia-
tion untenable. Original description date (31 December) 1878.
= Notropis hudsonius (Clinton)
Notropis santamariae Evermann and Goldsborough 1902
Evermann and Goldsborough 1902:147, fig. 2. TYPE LOCALITY: small pool near Lake Santa
BULLETIN FLORIDA STATE MUSEUM
Maria, Chihuahua, Mexico. HOLOTYPE: USNM 5(XX)2 (33.3), E. W. Nelson, September 1899.
PARATYPE: USNM 126485 (ex BCF 964) (1), paratopotype.
REMARKs.-Contreras (1975) recently elevated this form to a subspecies of N. lutrensis, and
considered it closely related to N. nltrensis fonnosus. Count for holotype: anal rays 8. Original
description date 3 May 1902.
= Notropis lutrensis saintamariae Evermann and Goldsborough
Notropis santarosaliae Meek 19)2
Meek 1902:85: pl. 19, lower fig. TYPE LOCALITY: Rio Nonava, Santa Rosalia, Chihuahua,
Mexico. HOLOTYPE: FMNH 3535 (37.2), S. E. Meek and F. E. Lutz, April-June 1901. PAHATYPES:
FMNH 59532 (17), paratopotypes.
REMARKS.-Miller (1976:11-12) synonyinized this form with N. jciczanimts. Counts for eight
paratypes (FMNH 59532): pharyngeal teeth 2,4-4,2 (2); anal rays 9 (6), 10 (2). Original descrip-
tion date (31) May 1902.
= Notropis jerneznus (Cope)
Photogenis scabriceps Cope 1868
Cope 1868a:166. TYPE LOCALITIES: tribes. of Kanawha (=New) R. (Sinking Cr., Walkers Cr.,
and others), and near Austinville, Virginia. SYNTYPES: ANSP 7588-7589 (2, 55.8-58.8) (Sinking
Cr.); ANSP 2726-2746 (22, 33.5-60.3) (Sinking Cr.); ANSP 29(04-2906 (3, 38.5-60.3) (Walkers
Cr.); ANSP 3039-3044 (6, 42.0-52.4) (unnamed creek in Wythe County, Virginia); ANSP 4160-
4161 (2, 31.5-37.9) (Sinking Cr.), E. D. Cope, August-September 1867.
REMARKS.-Although Fowler (1910:288) listed ANSP 7588 as "type," this cannot be consid-
ered a lectotype designation as he did not include an illustration nor an indication of standard
length. Fowler made no mention of the other syntypic series listed above, but data on the labels
in the jars show they should be so considered. Counts for ANSP 7588-7589: pharyngeal teeth
2,4-4,2, anal rays 8, predorsal scales 13, caudal-peduncle scales 5-2-5 (larger specimen); pharyn-
geal teeth 1,4-4,1, anal rays 8, predorsal scales 15, caudal-peduncle scales 5-2-5 (smaller speci-
men). Counts for ANSP 3039-3044: pharyngeal teeth 2,4-4,1 (1), 2,4-4,2 (5); anal rays 8 (4), 9 (2).
Counts for ANSP 4160-4161: pharyngeal teeth 2,4-4,2 (both); anal rays 8 (both). Counts for ANSP
2904-2906: pharyngeal teeth 2,4-? (1), ?-4,2 (1); anal rays 8 (3).
Jordan and Evermann (1896:268) erroneously synonymized this species with N. shmatlirdi.
Several early references (Jordan 1876b:289; Jordan 1882:850-851; et al.) indicate a much wider
geographic range for this species than it actually has. Original description date 11 May 1868.
= Notropis scabriceps (Cope)
Minnil.us scepticrts Jordan and Gilbert 1883
Jordan and Gilbert 1883:2(X)-201. TYPE LOAALITY: Saluda R., near Farr's Mill, W of Green-
ville, Greenville County, South Carolina. SYNTYPES: USNM 31081 (21, 44.0-57.4) (in two jars,
of which one contains one specimen), CAS-SU 2529 (1, 50.7), BMNH 1883.12.14.212 (1, 57.0),
BMNH 1880.1.21.83 (1, 46.5), MNHN A.1288 (5, 54.0-59.6), D. S. Jordan and A. W. Brayton,
summer 1877.
REMARKS.-Counts: anal rays 10 in two specimens counted (USNM 31081 and CAS-SU 2529).
Hubbs (1941) reviewed species. Original description date 1 April 1883.
= Notropis scepticus (Jordan and Gilbert)
Notropis scopiferus Eigenmann and Eigenmann 1893
Eigenmann and Eigenmann 1893a:153. TYPE LOC ALITIES: Winnipeg, Manitoba; Brandon,
Manitoba; Fort Qu'Appelle, Saskatchewan; Medicine Hat, Alberta. SYNTYPES: USNM 125160
(2, 60.4-66.5) (Fort Qu'Appelle), BMNH 1892.12.30.500-509 (38, 41.0-81.0) (Fort Qu'Appelle),
BMNH 1892.12.30.497-499 (3, 26.0-56.0) (Brandon), BMNH 1892.12.30.510-517 (8, 23.0-57.5)
(Medicine Hat), BMNH 1892.12.30.518-527 (23, 20.0-53.0) (Winnipeg), CU 1877 (2,50.0-63.0)
(Fort Qu'Appelle), C. H. Eigenmann, August or September 1892.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
REMARKS.-This species name has sometime been incorrectly spelled scopif r, as by Jordan
and Evermann (1896:291-292). Woolman (1896:351) was first to svnonvmize it with .V. huldsonius.
Original description date 4 Februarv 1893.
= Notropis hudsonius (Clinton)
Hybopsis scylla Cope 1871
Cope 1871:438. TYPE LocALITY: Red Cloud Cr., trib. to North Platte R. (Wyoming ?). TYPES:
None located.
REMARKS.-Jordan and Gilbert (1883:168) svnonvmized this forn with Cliola licolata
(Agassiz), which in turn is a probable senior synonym of X. stramuinus (Cope). Hobbs and
Greene (1928:377) restricted this to the subspecies missrien.sis. Inasmuch as Ilybopsis missurien-
sis and H. scylla were described in the same paper (Cope 1871:437-438), Hubbs and Greene's
choice of the former name qualifies them as first revisers. Bailey and Alluin (1962:65-68) con-
firmed the allocation of H. scylla to the synonvmv of N. straminm'eus missunriensis and provided
a clearer indication of the subspecies' range. Original description date (31 December) 1871.
= Notropis stramineus missuriensis (Cope)
Luxilus selene Jordan 1877
Jordan 1877e:60-61. TYPE LOCALITY: Lake Superior and tribes near Bavfield, Bayfield County,
Wisconsin. TYPEs: None located.
REMARKs.-Jordan (1877e), in his original description, said: "Many specimens in United States
National Museum: collected near Bayfield, Wis., by J. W. Milner," but he later (1885a:127) indi-
cated that only one specimen was involved, and at the same time synonymized se'lm'n with .o-
tropis hudsonius. Original description date (31 December) 1877.
= Notropis hudsoniius (Clinton)
Notropis semperasper Gilbert 1961
Gilbert 1961c:450-456, fig. 1. TYPE LOCA.LITY: Potts Cr., just above junction with Jackson
(James) R., at st. hwy. 18 bridge, near Covington, Alleghany County, Virginia. HOLOTYPE:
UMMZ 174815 (59.0), Carter R. Gilbert and John W. Motile, 14 June 1956. PARA'TYPES: UMMZ
174816 (4), UMMZ 175100 (10), paratopotypes; UMMZ 95199 and UMMZ 95458 (9), UMMZ
175076 (6), UMMZ 175118 (14), UMMZ 175133 (4), USNM 162868 (10).
REMARKS.-Jenkins and Bnrkhead (1975) reviewed species. Original description date 19 De-
cenmber 1961.
= Notropis sclmperasper Gilbert
Alburnops shumardi Girard 1857
Girard 1857:194. TYPE LOCALITY: Arkansas R., near Fort Smith, Arkansas. SYNTYPES: USNM
68 (one set of pharyngeal teeth only), George C. Shumard, 1853.
REMARKs.-Girard (1858:261-262; pl. 57, figs. 1-4) listed two original syntypes (USNM 68)
and illustrated one of them. Counts for set of existing pharyngeal teeth: 2,4-4,2. A. illecebrosus,
which was described on the same page of the same paper as A. shmnardi, was regarded as identi-
cal to that species by Jordan and Gilbert (1883:192-193), who, as first revisers, selected the name
shumardi. It now appears that Jordan and Gilbert's action, though correct, was based on the as-
sumption that both names apply to the species now called N. hoops (R. D. Suttkus, in litt.). Gil-
bert and Bailey (1962) reviewed the status and distribution of this species. Original description
date 25 April 1857.
= Notropis shnmardi (Girard)
Notropis signipinnis Bailey and Suttkus 1952
Bailey and Suttkus 1952:1-15, pls. 1 and 2B. TYPE LOCALITY: Maletts Spring Branch, trib. to
Rocky Cr. and Escatawpa R. (Pascagoula R. drainage), 6.6 mi SE of Lucedale, George County,
Mississippi. HOLOTYPE: UMMZ 157117(49.5), Marian K. and Reeve M. Bailey, 4 April 1948. PARA-
BULLETIN FLORIDA STATE MUSEUM
TYPES: UMMZ 155447 (erroneously listed as 155477 at one place in paper) (173 originally, now
153), USNM 163256 (ex UMMZ 155447) (20), TU 1133 (164), paratopotypes.
REMARKs.-Original description date 17 October 1952.
= Notropis signipinnis Bailey and Suttkus
AlIbrnelhls simus Cope 1875
Cope, in Cope and Yarrow 1875:649-650; pl. 31, figs. 2-2a. TYPE LOCALITY: Rio Grande, at
San Ildefonso, ca 10 mi E of Los Alamos, Santa Fe County, New Mexico. SYNTYPES: USNM 15800
(5, 25.5-43.0), USNM 16982 (72, 22.0-64.0), MCZ 359:33 (2, 49.0-70.0), E. D. Cope and H. C. Yar-
row, August 1874.
REMARKs.-Label on jar containing two MCZ specimens reads "pools of Rio Grande, Rio de
Acama, New Mexico." As "Rio de Acama" is not mentioned in the original description, the two
MCZ specimens possibly are not from the original syntypic series, and when a lectotype is desig-
nated one of the USNM syntypes should be selected.
Above syntypes all appear to represent the same species. Counts for USNM 158(X) (four
specimens): pharyngeal teeth 1,4-4,1 (3), 2,4-4,2 (1); anal rays 9 (4). Counts for MCZ syntypes:
pharyngeal teeth 2,4-4,2(1), 2,4-4.1 (1); anal rays 9 (1), 10 (1); lateral-line scales 37(1). Fins sharply
pointed. Miller (1976) discussed status of species. Original description date (31 December) 1875.
= Notropis simus (Cope)
Alburnus socius Cirard 1857
Girard 1857:193. TYPE LOCALITY: Liveoak Cr. (tributary to Pecos R.), ca 8 mi NE of Sheffield,
Crockett County, Texas. SYNTYPES: USNM 70 (17, 42.2-47.3, plus one set of pharyngeal arches in
bone collection) (also one specimen of N. colucellus earlier removed by R. D. Suttkus), USNM
39654 (2, 42.8-56.0), ANSP 2848 (ex USNM 70) (1, 48.7), MCZ 1683 (ex USNM 70) (2, 41.5-50.8),
John H. Clark 1851.
REMARKs.-The label in MCZ 1683 indicates three specimens as being present. Girard
(1859a:52; pl. 29, figs. 14-17) listed only 20 original syntypes and illustrated one of these. Jordan
(1885a:123) regarded this form as distinct from N. swaini (= N. amnabilis); and Evermann and
Kendall (1894:83), Jordan and Evermann (1896:292), and Jordan, Evermann, and Clark (1930:124)
also recognized it as valid. There appear to be no subsequent references to this species, nor any
indication of its being synonymized with any currently recognized species. Thus the present
synonymization of this form with N. amabilis appears to be the first time that this relationship
has been definitely established.
Counts for ANSP 2848: teeth 2,4-4,2; anal rays 9. Counts for MCZ 1683: teeth 2,4-4,2 (larger
specimen), 1,4-4,1 (smaller specimen); anal rays 9 in both; predorsal scales 15 and 16 (larger and
smaller specimens, respectively); caudal-peduncle scales 6-2-5 and 5-2-5 (larger and smaller speci-
mens, respectively); body-circumferential scales 12-2-11 (larger specimen). Counts for USNM
39654: teeth 1,4-4,1 (smaller specimen; teeth missing from larger individual); anal rays 9 in both;
caudal-peduncle scales 5-2-5 in both. Counts for USNM 70: teeth 1,4-4,1 (1), 2,4-4,1 (2): anal
rays 9 (3). All types in fair to poor condition. Original description date 25 April 1857.
= Notropis ,omabilis (Cirard)
Hyhopsis spectrnnculus Cope 1868
Cope 1868b:231; pl. 22, fig. 3. TYPE LOC(ALITY: Bear Cr., tribe. to Middle Fork of Holston
R., Smyth County, Virginia. SYNTYPES: ANSP 4363-4373 (11, 41.0-47.0), E. D. Cope, October
1867.
REMARKS.-Fowler (1910:275) listed ANSP 4363 as "type," but did not illustrate specimen,
give an indication of its length, nor segregate it from the remainder of the syntypic series. Under
the circumstances, this is not regarded as a lectotype designation. Counts: pharyngeal teeth 4-?
(1), anal rays 9 (1). Original description date 2 December 1868.
= Notropis spectrunculus (Cope)
Vol. 23, No. 1
1978
GILBERT: NOTROPIS TYPE CATALOGUE
Leuciscus spiloptcrus Cope 1868
Cope, in Ginther 1868:254. TYPE LOCALITY: St. Joseph R., Michigan (southwestern part of
state). LECTOTYPE: ANSP 22901 (46.0), E. D. Cope. Gibbs (1958:193) designated lectotype.
REMARKS.-The original description of this species (as Photogenis spilopterus) was long con-
sidered to date from Cope (1869:378), but I (Gilbert 1971a) recently showed that this was pre-
ceded by the earlier description (as Leuciscus spilopterus) in Giinther (1868:254). This species
was long regarded as either a junior synonym of N. tchipplei or subspecifically distinct. Hubbs
and Lagler (1943:78-79) first pointed out their specific distinctness. Fowler (1910: pl. 18, fig.
28) illustrated the "cotype." Gibbs (1958) reviewed species. Original description date 14 March
1868.
= Notropis spilopterus (Cope)
Notropis spilurus Gilbert and Swain 1885
See page 22.
Leuciscus spirlingulus Valenciennes 1844
Valenciennes, in Cuvier and Valenciennes 1844:321-323. TYPE LOCALITY: "New Jersey."
HOLOTYPE: MNHN 3892 (85.3). PARATYPES: MNHN 1709 (2).
REMARKS.-Jordan (1880a:225) was first to synonymize this form with Luxilus (= Notropis)
cornntus. Original description date (31) July 1844.
= Notropis cornutus (Mitchill)
Photogenis stigmaturus Jordan 1877
Jordan 1877c:337. TYPE LOCALITY: trib. of Etowah R. (probably Silver Cr.), near Rome, Floyd
County, Georgia. LECTOTYPE: USNM 17891 (84.4), D. S. Jordan, July 1876. Gibbs (1957:192)
designated lectotype. LECTOPARATYPES: USNM 163938 (ex USNM 17891) (10), USNM 20125
(1), ANSP 19820-19827 (8), MCZ 24371 (1), MNHN A.1283 (2).
REMARKS.-Syntypes came from tributaries of Coosa and Oostanaula rivers, in addition to
Etowah River. Gibbs (1957:192) restricted type locality by designating lectotype. Fowler
(1910: pl. 17, fig. 24) illustrated one of ANSP "cotypes." Bollman (1887:462) was first to call this
form a subspecies of N. venustus, though most authors (Jordan and Evermann 1896:255; Jordan,
Evermann, and Clark 1930:130) continued to regard it as a valid species. Bailey, Winn, and Smith
(1954:128) synonymized stigmaturus with N. cenustus. Gibbs (1957), in his review of N. cenustus,
recognized stigmaturus as a valid subspecies. Original description date (30) June 1877.
= Notropis renustus stigmaturus (Jordan)
Nototropis stilbius Jordan 1877
Jordan 1877c:343. TYPE LO(CALITIES: tribes. of Etowah, Oostanaula, and Coosa rivers, near
Rome, Floyd County, Georgia. SYNTYPES: USNM 17879 (2, 50.2-50.7) (Etowah R.) (not USNM
31132, as listed by Jordan and Evermann [1896:293]), ANSP 18701-18702 (53.0-59.0) (Etowah
R.), CAS-SU 2528 (3, 50.2-62.4) (Etowah R.), BMNH 1880.1.21.16 1, 60.0) ("Alabama R."),
MNHN A.1297 (1, 57.5) (Etowah R.), D. S. Jordan and C. H. Gilbert, summer 1876.
REMARKS.- Condition of syntypes fair (ANSP) to good (others). Counts for ANSP 18701: larger
specimen with pharyngeal teeth 2,4-4,2, anal rays 10; CAS-SU 2528: pharyngeal teeth 2,4-4,2
(1), 1,4- (1), anal rays 10 (3); USNM 17879: anal rays 10 in both. Fowler (1910: pl. 20, fig. 46) illus-
trated larger ANSP "cotype." Original description date (30) June 1877.
= Notropis stilbius (Jordan)
Notropis stone Fowler 1921
Fowler 1921:391-393, fig. TYPE LOCALITY: Pocataligo R., near Manning, Clarendon County,
South Carolina. Moore (1957:129) questioned the type locality because, according to him, recent
collections from this river system have failed to yield the species. HOLOTYPE: ANSP 50118 (ca
30.0), Witmer Stone, May 1917. PARATYPES: ANSP 50119-50121 (3), paratopotypes.
REMARKS.-Types not located by me at ANSP, but believed by J. E. Bohlke (pers. comm.)
BULLETIN FLORIDA STATE MUSEUM
not to be lost. Bailey and Suttkus (1952:13) and Suttkus (1955:98) regarded this form as a distinct
species. Moore (1957:129) seems to have been the first to note its conspecificity with N. hyp-
selopterus. It was not regarded as a distinct species in either the 1960 or 1970 AFS checklists
(Bailey et al. 1960, 1970). Original description date 4 March 1921.
= Notropis hypselopterus stonei Fowler
Hybognathus stramineus Cope 1865
Cope 1865a:283. TYPE LOCALITY: Detroit R., Grosse Isle, Michigan. LECTOTYPE: ANSP 4131
(51.0), Prof. Manly Miles. Fowler (1910: pl. 15, fig. 5) designated lectotype by illustrating "type"
(ANSP 4131); calculated length 53.3 mm SL. LECTOPARATYPES: ANSP 413241136 (5), paratopo-
types.
REMARKs.-This species was often erroneously called N. blennius in the early literature;
Fowler (1910:274-276) first demonstrated, and Hubbs (1926:42-43) subsequently confirmed, that
blennius is a distinct species. The form stramineus was regarded either as a subspecies (Jordan
1885b:811; Hubbs 1926:37, 43) or a junior synonym of N. delicious until Suttkus (1958) showed
that the lectotype of Moniana deliciosa earlier designated by Clark Hubbs (1954a) represents the
species now called N. texanus (Girard). It should be noted that none of the extant syntypes of
M. deliciosa are N. stramineus, although some specimens of this species may have been present
in the original series. Bailey and Allum (1962:65) showed the subspecific separation of delicious
and stramineus proposed by Hubbs (1926:37, 43) to be invalid, but at the same time they demon-
strated (1962:64-68) the subspecific distinctness of stramineus and missuriensis. N. stramineus
and N. volucellus also were variously regarded as subspecies (Jordan 1885b:811), synonyms, or as
distinct species in the early literature; Hubbs and Greene (1928:375) were the first to demonstrate
conclusively their specific distinctness. Alburnus lineolntus Agassiz 1863 likely is a senior syn-
onym of N. stramineus, but inasmuch as no unquestioned types appear to be extant, this cannot
be proved conclusively. One of the two extant syntypes of Cyprinella ludibunda Girard 1857
represents N. stramineus; the other syntype is N. volucellus (Cope 1865). Original description
date 13 February 1865.
= Notropis stramineus stramineus (Cope)
Cyprinella suavis Girard 1857
Girard 1857:197. TYPE LOCALITY: near San Antonio, Texas. SYNTYPES: USNM 138 (one set of
pharyngeal teeth in osteological collection), ANSP 2769-2770 (ex USNM 138) (2, 26.5-34.5),
MCZ 1694 (ex USNM 138) (2, 28.3-34.5), Dr. C. B. Kennerly, 1853. Two other syntypes (USNM
138) are supposedly present in the USNM, but these cannot presently be located.
present in the USNM, but these cannot presently be located.
REMARKS.-Girard (1858:268) listed 24 original syntypes (USNM 138) but provided no illustra-
tion. Counts for USNM 138: pharyngeal teeth 4-4. Counts for ANSP 2769-2770: pharyngeal teeth
4-4 (1), 4-? (1); anal rays 9 in both. Counts for MCZ 1694: pharyngeal teeth 4-4 in both; anal rays
9 in both. Jordan (1885a:124) was first to synonymize this form with N. lutrensis. Carl L. Hubbs
(1954:292) and Clark Hubbs (1954b:282-283) called it a subspecies of N. lutrensis. Original de-
scription date 25 April 1857.
= Notropis lutrensis (Baird and Girard)
Hybopsis harper subterranea Hubbs and Crowe 1956
Hubhs and Crowe 1956:3-5. TYPE LOCALITY: Jerome Sink, S of High Springs, Alachua County,
Florida. HOLOTYPE: UMMZ 133551 (33.8), R. R. Miller, B. W. Walker, and N. A. Marshall, 4
April 1941. PARATYPES: UMMZ 197700 (99) (not 111 as stated in original description), paratopo-
types.
REMARKs.-Howell (1960) demonstrated that this form is not a valid subspecies. Original
description date 24 August 1956.
= Notropis harper Fowler
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
Notropis swaini Jordan and Gilbert 1885
Jordan and Gilbert, in Jordan 1885a:123. TYPE LOCALITY: San Felipe Cr., near Del Rio, Val
Verde County, Texas. SYNTYPES: USNM 36529 (ex USNM 69) (1, 40.2), ANSP 3246 (ex USNM
69) (1, 36.5), MCZ 1682 (ex USNM 69) (2, 35.0-39.5), John H. Clark, 1851. (Label in jar of MCZ
types says "1853").
REMARKs.-Jordan and Gilbert (in Jordan 1885a:123) proposed this as a substitute name for
Alburnus megalops Girard 1857, which is preoccupied in the genus Notropis by Cyprinus mega-
lops Rafinesque 1817. Girard (1859a: pl. 29, figs. 1-4) listed 20 original syntypes of A. megalops
(USNM 69) and illustrated one of these. Hubbs, Kuehne, and Ball (1953:226) indicated that rec-
ords of N. swaini from the Guadelupe River system are based on N. amabilis, but did not formally
synonymize the two forms in that paper. For other remarks see account of A. megalops. Original
description date 19 June 1885.
= Notropis amabilis (Girard)
Notropis altipinnis tarensis Hubbs and Ranev 1948
Hubbs and Ranev 1948:13-14. TYPE LOCALITY: small stream, 4.5 mi E of Surl, on US hwv.
158, Person County, North Carolina. HOLOTYPE: UMMZ 136148 (42.2), E. C. Ranev, E. A. Lach-
ner, and R. A. Pfeiffer, 2 April 1941. PAHATYPES: UMMZ 138526 (4), CU 10570 (25), CU 9840
(1), paratopotypes.
REMARKS.-This subspecies and others described in the same paper (Hubbs and Raney 1948)
are not generally regarded as valid, although they have never been formally synonymized. Orig-
inal description date 26 April 1948.
= Notropis altipinnis (Cope)
Leuciscus telescopes Cope 1868
Cope, in Ginther 1868:252-253. TYPE LOC(ALITY: Holston R. and tribs., Virginia. LECTOTYPE:
ANSP 2157 (67.0), E. D. Cope, October 1867. Gilbert (1969:487) designated lectotype. LECTO-
PARATYPES: ANSP 2158-2254 (97) (Holston R.), ANSP 2330-2335 (6) (tribs. of Holston R.), MCZ
25160 (1), BMNH 1868.1.10.18 (7), MNHN 4855 (1).
REMARKS.-This species was long considered to have been described originally by Cope
(1868a) as Photogenis telescopes, but I (Gilbert 1971a) showed Cope's description was preceded
by the one appearing in Giinther (1868). N. telescopus was regarded as a valid species until Kuhne
(1939:49) downgraded it to a subspecies of N. ariommus. I (Gilbert 1969) later re-elevated it to a
full species. Fowler (1910:289) listed ANSP 2157 as the "type" number, but this cannot be con-
sidered a lectotype designation because he did not illustrate a specimen, segregate one from the
remaining syntypes, nor provide an indication of body length. Counts for lectotype: pharyngeal
teeth 2,4-4,2: anal rays 10. Gilbert (1969) reviewed species. Original description date 10 March
1868.
= .otropis telhscopus (Cope)
Cyprinella texana Girard 1857
Girard 1857:198. TYPE LO(:ALITY: Salado Cr., just E of San Antonio, Bexar County, Texas.
LECTOTYPE: USNM 128 (46.3), John H. Clark, 1851. Pharyngeal arches from lectotype previously
removed and now in USNM osteological collection, also bearing catalogue number USNM 128
(Suttkus 1958:316). Clark Hubbs (1954a) designated lectotype. LECTOPARATYPES: USNM 162721
(3), paratopotypes. The original syntypes (USNM 162722 [2], ANSP 3010 [1], and MCZ 1693
[1]) were identified by Suttkus (1958:308) as Notropis rolucellus.
REMARKs.-Girard (1859:55; pl. 31, figs. 9-12) listed 10 original syntypes of C. texana and il-
lustrated one of these. Nine syntypes are from Salado Creek, Texas, and the other from Turkey
Creek, Texas. Clark Hubbs (1954a) erroneously considered the lectotype and three lectopara-
types of C. texana to be N. venustus; these were subsequently reidentified by Suttkus (1958).
Jordan (1885a:124) synonymized C. texana with N. notatus (= N. venustus), and it was so con-
1978
BULLETIN FLORIDA STATE MUSEUM
sidered prior to publication of Suttkus's (1958) paper. N. texanus, as now recognized, was called
N. roseus before 1958. Counts for lectotype of C. texana: pharyngeal teeth 2,44,2; anal rays 6
(aberrant count for species, which usually has 7 rays). Counts for lectoparatypes: pharyngeal
teeth 2,4-4,1 (2), 2,4-4,2 (1); anal rays 7 (3). Swift (1970) reviewed species. Original description
date 25 April 1857.
= Notropis texanus (Girard)
Cliola (Hybopsis) topeka Gilbert 1884
Gilbert 1884c:13. TYPE LOCALITY: Shunganunga Cr., trib. to Kansas R., Shawnee County,
Kansas. SYNTYPES: USNM 36609 (4, 47.0-49.6), MCZ 35931 (1, 49.8), F. W. Cragin, 1884.
REMARKs.-Gilbert (1885:513) subsequently republished original description verbatim "for
convenience of reference." Minckley and Cross (1959) reviewed species. Counts for USNM syn-
types: pharyngeal teeth 4-4 (1); anal rays 7 (1). Counts for MCZ 35931: pharyngeal teeth 4-4;
anal rays 7. Condition of syntypes good. Original description date (30) September 1884.
= Notropis topeka (Gilbert)
Codoma trichroistia Jordan and Gilbert 1878
Jordan and Gilbert, in Jordan and Brayton 1878:50-51. TYPE LOCALITY: tributary of Etowah
R., Georgia. (Probably near Rome, Floyd County, Georgia). LECTOTYPE: USNM 31131 (54.4),
D. S. Jordan and C. H. Gilbert, summer 1877. Howell and Williams (1971:63) designated lecto-
type. LECTOPARATYPES: USNM 163956 (ex USNM 31131) (6), paratopotypes; MCZ 24393 (1),
MNHN A.1280 (1).
REMARKS.-Gibbs (1955) reviewed species. Howell and Williams (1971) mapped distribution,
illustrated nuptial tubercle pattern, presented meristic data, and discussed differences between
this species and the closely related N. gibbsi. Original description date (31 December) 1878.
= Notropis trichroistius (Jordan and Gilbert)
Moniana tristis Girard 1857
Girard 1857:201. TYPE LOCALITY: Unknown. TYPES: None located. Girard (1858:278) listed
five specimens (USNM 93), supposedly collected in 1854 by Dr. Creutzfeldt, who had been killed
the preceding October.
REMARKs.-Inasmuch as Girard (1857, 1858) neither illustrated nor adequately described
this form, no types apparently remain, and the provenance of the types remains in doubt; a posi-
tive identification cannot be made. Jordan and Evermann (1896:272) regarded it as a question-
able synonym of N. proserpinus, hut this is highly improbable as the route followed by the Gunni-
son party (which collected the specimens) was outside the range of that species. Furthermore
the combined characters of a slender body, a long caudal peduncle of nearly uniform depth, and
seven anal rays further eliminate N. proserpinus from consideration. This form most likely is a
species of Notropis, but even this cannot be determined categorically. Original description date
25 April 1857.
= Not definitely identifiable.
Cyprinus tricittatus Rafinesque 1820
Rafinesque 1820a:6. TYPE LOCALITY: Fish and Wallkill creeks, New York. TYPES: None lo-
cated.
REMARKS.-I (Gilbert 1964:141) synonymized this form with N. cornutus. Original descrip-
tion date 1 March 1820.
= Notropis cornutus (Mitchill)
Notropis tropics Hubbs and Miller 1975
Hubbs and Miller 1975:121-131, figs. 1-2. TYPE LOCALITY: Rio Guayelejo, at Llera, Tamall-
lipas, Mexico. HOLOTYPE: UMMZ 193804 (30.5), Rezneat M. Darnell et al, 20 June 1968. PARA-
TYPES: UMMZ 192897 (49), paratopotypes; UMMZ 97409 (1), UMMZ 97410 (1), UMMZ 97411
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
(1), UMMZ 97412 (2), UMMZ 124324 (1), UMMZ 124344 (1), UMMZ 161656 (1), UMMZ 162109
(3), UMMZ 164710 (28), UMMZ 169498 (1), UMMZ 18()033 (46), UMMZ 193806 (2).
REMAHKs.--Original description date 15 May 1975.
= Notropis tropics Hubbs and Miller
Plargyrus typicus Cirard 1857
Girard 1857:196. TYPE LOCA.LTY: "Kentucky." TYPES: None located.
REMARKS.-This name was proposed to replace Rutilus plargyrus, which was earlier described
by Rafinesque (1820b:50-51). According to Girard (1857:195), "Since Rafinesque's genera are to
be restored, his genus Plargyrns is to take precedence over the genus Hypsolepis of more modern
coining. The name of Plargyruis was provided for in the Ichthyologia Ohicnsis to replace that of
Rutilus in the eventuality that Cyprinus rutilus of Europe, which was the type of the genus
Rutilus, should prove generically distinct from Rutilus plargyrus and similar American species,
and which is the case." Original description date 25 April 1857.
= Notropis chrysocephalus chrysocephalis (Rafinesque)
All/tirnus roibratilis Girard 1857
Girard 1857:193. TYPE LOCALITY: Sugarloaf Cr., trib. to Poteau R., about 20-25 mi S of Fort
Smith, Arkansas, vicinity of Poteau, Oklahoma. Locality is very near the Oklahoma-Arkansas
state line, and could either be in Le Flore County, Oklahoma or Sebastian County, Arkansas
(See REMARKs below). LECTOTYPE: USNM 73 (56.4), H. B. Mollhausen, probably 25 July 185:3.
Snelson and Pflieger (1975:233) designated lectotype. LECTOPARATYPES: USNM 203328 (ex
USNM 73) (39) (seven additional original syntypes, representing two other species, removed and
recatalogued), USNM 2678 (ex USNM 73) (one pair of pharyngeal arches in osteological collec-
tion), MCZ 1786 (ex USNM 7:3) (2) (not seen bv me).
REMARKS.-The type locality was said by Girard (1857:193: 1858:260) to be in Arkansas, and
was so listed by Snelson and Pflieger (1975:233), but Gibbs (1963:513) said that this actually was
in Oklahoma. Girard (1858:260) listed only 24 original svntypes. Snelson and Pflieger (1975) re-
viewed species. Jordan (1885a:122) synonvmized Luxilus iuicidus Girard 1857 with N. umrbratilis,
and as first reviser selected the latter name. Original description date 25 April 1857.
= Notropis umbratilis irnbratilis (Girard)
Notropis urnbrifer lHay 1887
Hay 1887:245. TYPE Lc( \LITY: Saline R., about 5 mi N of \a Keeney, Trego Count\.
Kansas; and Solomon River, at Beloit, Mitchell County, Kansas. TYPES: None designated. One
specimen was listed from Beloit and four from Wa Keenev. These were not located Ib me,
although they may be present in the general (non-type) collection of the USNM under a different
name. The specimens were collected by O. P. Hay and M. J. Thompson in July 1885.
REMARKS.-The five specimens upon which this name was based were initially identified by
Hay (1887:244-245) as N. macrostomna (Girard), with mnibrifer being proposed in the same paper
as a substitute name "in case future investigations should prove that these specimens are not to
he referred to Girard's species." Hay also stated "at first view these have a striking resemblance
to Notropis topeka (Gilbert." Cross (1967:125) included Hay's record of N. macrostoma in the
synonymy of N. hltrensis, thus in effect synonymizing N. umbrifer with that species. This pre-
sumably was based largely on the pharyngeal-tooth and anal-ray counts (1,4-4,1 and 9, respec-
tively) listed by Hay in his description as only two species of Kansas Cyprinidae (N. lutrensis
and N. c(aniurus) have this combination of counts, and of these only the former is known to occur
in the northwestern part of the state. Further study of the description, considered with other
factors, raises serious doubts that Cross' interpretation is correct. In the first place, Hay was very
familiar with N. lutrensis, and he mentioned collecting numerous specimens at both the Beloit
and Wa Keeney localities. It seems unlikely, therefore, that he would have misidentified speci-
mens of such an abundant and widespread species. Beyond this, a number of characters listed
BULLETIN FLORIDA STATE MUSEUM
in the description of "N. mnacrostomna-utmbrifer" are not concordant with those of N. lutrensis.
These include (a) an extremely oblique mouth (mouth moderately oblique in N. lutrensis); (b)
strongly serrated edges on the pharyngeal teeth (edges slightly crenulate in N. lutrensis); (c) a
dusky to dark stripe along the sides of the body and caudal peduncle (not present in N. lutrensis);
(d) snout shorter than eye (longer than eye in N. lutrensis); and (e) eye contained three times in
length of head (contained at least four times in head in lutrensis).
Assuming that the above interpretation is correct, it seems that N. umbrifrr is (a) a valid spe-
cies that is either extinct or, if living, has not been collected since 1885; or (b) is based on a hybrid,
one or both parents of which may no longer occur in northwestern Kansas or perhaps elsewhere
in the state (see Hubbs 1951b). Of these, the latter possibility is much more likely. In view of the
alleged similarity in appearance to N. topeka (a species having 4-4 teeth and 7 anal rays), that
species is suggested as one possible parent. If so, the other parent would logically be a species
having 2,4-4,2 teeth and 10 or 11 anal rays, such as N. umbratilis, N. rubellus, or N. atherinoides;
none of these species occur today in northwestern Kansas (Cross 1967:102, 104, 108), but may
have occurred there in the past. Until some or all of Hay's original specimens are located, the
identity of the parental forms (if, in fact, N. mibrifer is of hybrid origin) can only be surmised.
Inasmuch as a number of other series of specimens collected by Hay and reported upon by him
in his 1887 paper are still extant, there is a good chance that the types of Notropis minbrifer will
eventually be found, most likely in the USNM general (non-type) collection. Original descrip-
tion 2 July 1887.
= Identity uncertain. Most likely based on a hybrid, one parent of which may le Notropis
topeka.
Cyprinella uimbrosa (Girard 1857
Cirard 1857:197. TYPE LOCALITY: Coal Cr., trib. to South Fk. of Canadian R., N of McAlester,
Pittsburg County, Oklahoma. SYNTYPES: USNM 133 (6, 51.8-58.8), USNM 134 (one set of pharyn-
geal arches in bone collection), ANSP 2830 (1, 55.7), MCZ 18(0 (1, 53.7), H. B. Mollhausen, 7-8
August 1853.
REMAIKs.-Types in relatively good condition. Counts: USNM 134 pharyngeall teeth 1,4-
4,0; lesser tooth in left arch represented by a small hole); ANSP 2830 pharyngeall teeth 1,4-?;
anal rays 10); MCZ 18(X) pharyngeall teeth ?-4; anal rays 9. Jordan (1885a:123) was first to synon-
vmize this form with N. bubalinus (= N. lutrensis). Original description date 25 April 1857.
= Notropis lutrensis (Baird and Girard)
Notropis unirersitatis Evermann and Cockerell 1909
Evermann and Cockerell 1909:187. TYPE LOCALITY: Boulder Cr. (trib. to South Platte R.),
Boulder, Boulder County, Colorado. HOLOTYPE: USNM 64151 (51.0), T. D. A. Cockerell, 1907.
REMARKS.- Hubbs (1926:46) was first to synonymize this form with N. cornutus frontalis
(= N. cornutus). Original description date 8 December 1909.
= Notropis cornutus (Mitchill)
Notropis iranoscopus Suttkus 1959
Suttkus 1959:7-11, fig. 1. TYPE LOCALITY: Cahaba River, 2.2 mi N of Centreville, Bibb County,
Alabama. HOLOTYPE: TU 15289 (44.7), R. D. Suttkus, R. J. and H. C. Miller, and J. DeAbate,
17 March 1957. PARATYPES: TU 18551 (18), paratopotypes; TU 18569 (2), TU 18570 (8), TU
18674 (8), TU 19410 (21), UMMZ 171147 (2), UMMZ 171747 (12), UAIC 404 (3), UAIC 408 (1).
REMARKS.-Original description date 17 April 1959.
= Notropis nranoscopus Suttkus
Clioln urostigma Jordan and Meek 1884
Jordan and Meek 1884:474-477. TYPE LO( ALITY: San Saba R. (trib. to Colorado R.), at Fort
McKavit (= McKavett), ca. 45 air mi SE of San Angelo, Menard County, Texas. LECTOTYPE:
USNM 20446 (76.3), W. W. Anderson. Gibbs (1957:181) designated lectotype. LECTOPARATYPES:
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
USNM 163952 (8), MCZ 35903 (1, completely disintegrated), paratopotypes; USNM 17812 (7),
MCZ 35948 (1).
REMARKS.-Type locality restricted by Gibbs (1957:181) by lectotype designation. Other orig-
inal syntypes (USNM 17812 and MCZ 35948) are from Clear Creek, Waller County. Texas, col-
lected by Kumlein and Earll. Jordan (1885a:124) was first to synonymize C(lioa urrostigma with
N. cercostigma, and Gibbs (1957:179-181) synonymized the former with .V. trenustiis r'niiistus.
Original description date 23 October 1884.
= Votropis tenuistis renustus (Girard)
Cyprinella Ienutsta Girard 1857
Girard 1857:198. TYPE LOCALITY: Rio Sabinal, at Sabinal, Uvalde County, Texas. LE'(TOTYPE:
ANSP 3035 (ex USNM 140) (49.5), C. B. Kennerly, 1853. Gibbs (1957:180) designated lectotype.
LECTOPARATYPES: ANSP 3036 (ex USNM 140) (1), USNM 140 (one pair of pharyngeal arches in
bone collection), MCZ 1695 (ex USNM 140) (2), paratopotypes.
REMARKs.-Girard (1859a:55; pl. 31, figs. 1-4) listed 15 original syntypes and illustrated one of
them. Condition of types fair to poor (bodies soft), but pertinent characters evident. Counts: anal
rays 8 (ANSP 3035 and ANSP 3036); pharyngeal teeth 1,4-4,1 (USNM 140). Moore (1952:5), fol-
lowing then existing rules regarding line priority, used (without comment) the name Notropis
notatus instead of N, cenustus, both forms having been described in the same paper. Clark
Hubbs (1954b:280), as first reviser, chose the name renustus over notatus. Gibbs (1957) reviewed
species. Original description date 25 April 1857.
= Notropis venustus vennstus (Girard)
Leuciscus rittatus DeKav 1842
DeKay 1842:212; pl. 34, fig. 108. TYPE LOCALITY: Chittenonda Cr. (trib. to Mohawk R.), New
York. TYPEs: None located.
REMARKS.-Jordan and Evermann (1896:282) were first to synonymize this form with X. cor-
nutus. Original description date (31 December) 1842.
= Notropis cornutus (Mitchill)
Codonma cittata Girard 1857
Girard 1857:195. TYPE LOCALITY: Valley of Mexico. SYNTYPES: USNM 92 (one set of pharyn-
geal arches in osteological collection), John Potts, 1855.
REMARKS.-Girard (1859a:53-54: pl. 29, figs. 18-21) listed 10 original svntypes (USNM 92)
and illustrated one of them. Counts for the one set of extant pharyngeal arches (USNM 92):4-4.
Jordan (1885a:123) referred this species to the genus Notropis. Jordan and Evermann (1896:258)
were first to point out that it was preoccupied in Notropis by Leuciscuns rittatus DeKay, and thus
used the name N. aztecns Woolman for the species. Original description date 25 April 1857.
= Notropis sallei (Gunther)
Hyhognathris colucellus Cope 1865
Cope 1865a:283-284. TYPE LO):ALITY: Detroit R., at Grosse Ile, Wavne County, Michigan.
TYPEs: None located. Fowler (1910, 1918) made no mention of these being in the ANSP collec-
tion.
REMARKS.--N. straimines and V. rolucrellsu were variouslv regarded as subspecies (Jordan
1885b:811), synonyms, or as distinct species in the early literature; Hubbs and Greene (1928:375)
were the first to demonstrate conclusively their specific distinctness. At least two undescribed
species are presently going under the species name oliucellus. John Ramsey (unpubl. ms) is cur-
rently reviewing species. Original description date 1:3 February 1865.
= .Votrorpis rolucellus (Cope)
Notropis u'acrcamanus Fowler 1942
Fowler 1942:6-8, figs. 7-8. TYPE LO(ALITY: Lake Waccamaw, Columbus County, North Caro-
BULLETIN FLORIDA STATE MUSEUM
Vol. 23, No. 1
lina. HOLOTYPE: ANSP 69972 (56.5), Francis Harper, 22 May 1939. PARATYPES: ANSP 69973-
69978 (5), paratopotypes.
REMARKS.-Frey (1951:16-18) demonstrated that N. petersoni and N. wiarclanuiis are syn-
onyms. Both forms were described by Fowler (1942) in the same paper, and Frey (1951), as first
reviser, selected the name petersoni over waccamanus. Six paratypes presumably were present
originally. Counts for holotype: pharyngeal teeth 2,4-4,2; anal rays 7. Original description date
16 September 1942.
= Notropis petersoni Fowler
Notropis welaka Evermann and Kendall 1898
Evermann and Kendall 1898:126-127; pl. 6, fig. 2. TYPE LOCALITY: St. Johns R., near Welaka.
Putnam County, Florida. HOLOTYPE: USNM 48786 (36.3), W. C. Kendall, 19 March 1897. PARA-
TYPES: USNM 48785 (6), USNM 127014 (ex USFC 529) (16), UMMZ 172294 (ex USNM 127014)
(2), CAS-SU 9053 (14) (plus one specimen of N. petersoni removed), paratopotypes.
REMARKS.-CAS-SU 9053 erroneously listed as SU 5773 in original description. Pharyngeal
teeth of holotype now in USNM osteological collection. The pharyngeal tooth count was errone-
ously listed in the original description as 4-4, but it was subsequently found, upon re-examination
of the holotype, to be 1,4-4,1. This species was not recognized as valid for many years after its
original description, and Bailey, Winn, and Smith (1954:129) synonymized it with Notropis macru-
latus. Cook (1959:126-127) apparently was the first to resurrect N. wiehka as a valid species. Orig-
inal description date 8 February 1898.
= Notropis welaka Evermann and Kendall
Cyprinella whipplii Girard 1857 emendedd spelling Cyprinella whipplei)
Girard 1857:198. TYPE LOCALITY: Sugarloaf Cr., trib. to Poteau R., about 20-25 mi S of Fort
Smith, Arkansas, vicinity of Poteau, Oklahoma. Locality is very near the Oklahoma-Arkansas
state line, and could either be in Le Flore County, Oklahoma, or Sebastian County, Arkansas
(see REMARKS under Alburnus umbratilis). HOLOTYPE: USNM 137 (58.7), H. B. Mollhausen, prob-
ably 25 July 1853. Gibbs (1963:513) designated this specimen as "lectotvpe," but inasmuch as it
apparently was the only specimen collected (Girard 1858:270), it should properly be considered
the holotype.
REMARKS.-Girard (1858:270; pl. 58, figs. 6-10) listed one type specimen, which he illustrated.
Right pharyngeal arch of holotype in USNM osteological collection; count -4,1. N. whipplei was
long regarded as either a senior synonym of N. spilopterns or subspecifically distinct. Hubbs and
Lagler (1943:78-79) were the first to point out their specific distinctness. Gibbs (1963) reviewed
species. Original description date 25 April 1857.
= Notropis whipplei (Girard)
Notropis altipinnis white Hubbs and Ranev 1948
Hubbs and Raney 1948:15-17. TYPE LOCALITY: Morgan Cr. (trib. to Cape Fear R.), near
Chapel Hill, 1 mi upstream from bridge on road to Pittshoro, Orange County, North Carolina.
HOLOTYPE: UMMZ 128914 (37.1), A. McLaren White, 3 June 1933. PARATYPES: UMMZ 138528
(26), CU 8535 (21), paratopotypes, UMMZ 138529 (81), UMMZ 139742 (46), UMMZ 145266
(10), CU 3228 (2), CU 3537 (9), CU 1(X)75 (50). An undetermined number of paratypes were
designated in the original description, including 49 paratopotypes. Not all of these can be ac-
counted for.
REMARKS.-This form and others described in the same paper are not generally regarded as
valid, although they have never been formally synonymized. Original description date 26 April
1948.
= Notropis altipinnis (Cope)
Notropis colucellus wickliffi Trautman 1931
Trautman 1931:468-474, fig. 1. TYPE LOCALITY: Mouth of Miami R. at confluence with Ohio
GILBERT: NOTROPIS TYPE CATALOGUE
R., extreme SW Miami Twp., Hamilton County, Ohio. HOLOTYPE: UMMZ 92411 (48.0), M. B.
Trautman and R. B. Foster, 25 May 1931. PARATYPES: UMMZ 92412 (13), paratopotypes; UMMZ
76791 (1), UMMZ 76806 (30), UMMZ 78090 (12), UMMZ 78146 (1), UMMZ 78166 (27), UMMZ
78204 (4), UMMZ 78255 (5), UMMZ 78284 (6), UMMZ 81321 (13), UMMZ 86010 (2), UMMZ
87516 (17), UMMZ 87600 (4), UMMZ 91548 (1), USNM 93427 (ex UMMZ 76806) (1), USNM
117571 (ex UMMZ 76806) (3), OSM 10409 (9), OSM 10410 (109), OSM 10423 (31).
REMARKS.- Although this has been regarded as a valid subspecies since its original description,
J. S. Ramsey now thinks (pers. comm.) that it deserves full species status. Original description
date (30) November 1931.
= Notropis wickliffi Trautman
Notropis williamii Fowler 1945
Fowler 1945:236-237, figs. 117-119. TYPE LOCALITY: Brier Cr. (trib. to Savannah R.), ca 20
mi SW of Augusta, Richmond County, Georgia. HOLOTYPE: ANSP 70702 (60.0), William B. Har-
per, 5 July 1933. PARATYPES: ANSP 70703-70704 (2), paratopotypes.
REMARKs.-Suttkus and Raney (1955a:84) synonymized this form with N. petersoni. Counts
for holotype: pharyngeal teeth 2,44,2; anal rays 7. Original description date 25 October 1945.
= Notropis petersoni Fowler
Hybopsis winchelli Girard 1857
See pages 21, 92.
Notropis altipinnis wrighti Hubbs and Ranev 1948
Hubbs and Raney 1948:11-13. TYPE LOCALITY: Steele Cr. (trib. to Santee R.), Rock Hill, York
County, South Carolina. HOLOTYPE: UMMZ 94551 (30.2), Donald Ameel, 11 November 1931.
PARATYPES: UMMZ 94552 (3), paratopotypes; CU 10577 (23). Original description listed 10 other
paratypes, but their disposition is unknown.
REMARKS.-This form and others described in the same paper are not generally regarded as
valid, although they have never been formally synonymized. Original description date 26 April
1948.
= Notropis altipinnis (Cope)
Hyhopsis xaenocephalus Jordan 1877
Jordan 1877c:334-335. TYPE LOCALITY: Trib. of Etowah R., near Rome, Floyd County, Geor-
gia. LECTOTYPE: USNM 20116 (50.1), D. S. Jordan and C. H. Gilbert, summer 1876. Suttkus and
Raney (1955c:30) designated lectotype. LECTOPARATYPES: USNM 17886 (2), paratopotypes; CAS-
SU 3875 (1), ANSP 19828-19829 (2), MCZ 24394 (1). Remaining seven ANSP types represent
two other species, and have been recatalogued under those names.
REMARKS.-Fowler (1910: pl. 20, fig. 42) illustrated one of ANSP "cotypes." The species name
xaenocephalus first appeared as a nomen nudum in Jordan and Copeland (1877:151). Although
recognized as valid since its original description, N. xaenocephalus was frequently confused with
other forms of Notropis, particularly N. texanus and N. petersoni, and many references to "N.
xaenocephalus" are based on misidentifications involving those species. Bailey, Winn, and Smith
(1954:125-126) were the first to diagnose N. xaenocephalus clearly and define its range, and this
was further amplified by Suttkus and Raney (1955c). Counts for holotype: pharyngeal teeth 2,4-
4,2; anal rays 7. Original description date (30) June 1877.
= Notropis xaenocephalus (Jordan)
Minnilus xaenurus Jordan 1877
Jordan 1877b:79. TYPE LOCALITY: S. Fk. of Ocmulgee R., at Flat Shoals, DeKalb County,
Georgia. LECTOTYPE: USNM 17862 (66.0), D. S. Jordan and C. H. Gilbert, summer 1876. Lecto-
type herein designated by R. H. Gibbs, Jr. LECTOPARATYPES: USNM 163937 (ex USNM 17862)
(4), CAS-SU 1996 (1), MCZ 24365 (1), BMNH 1880.1.21.82 (1), MNHN A.1313 (2), paratopotypes.
REMARKS.-Condition of types good for old specimens. Counts for CAS-SU 1996: pharyngeal
BULLETIN FLORIDA STATE MUSEUM
teeth 1,4-; anal rays 10. Counts for MCZ 24365: anal rays 10. Gibbs (1955) reviewed species. Orig-
inal description date 17 April 1877.
= Notropis xaenurus (Jordan)
Notropis xanthicara Minckley and Lytle 1969
Minckley and Lytle 1969:491-502, fig. 1. TYPE LOCALITY: Rio Puente Colorado, 8.5 km S and
0.7 km W of Cuatro Cienegas, Coahuila, Mexico. HOLOTYPE: UMMZ 188782 (45.0), R. R. Miller
and family, C. L. Hubbs, D. R. Tindall, and W. L. Minckley, 6 April 1961. PARATYPES: UMMZ
179834 (11), ASU 969 (8), paratopotypes; ASU 2316 (185), UMMZ 179202 (46), UANL 703 (6),
UANL 709 (105), ASU 3728 (68), KU 7404 (72), UMMZ 179860 (53).
REMARKs.-Lytle (1972) reviewed species. Original description date 17 November 1969.
= Notropis xanthicara Minckley and Lytle
Ceratichthys zanemus Jordan and Brayton 1878
See pages 21, 92.
Alburnus zonatus Agassiz 1863
Agassiz, in Putnam 1863:9. TYPE LOCALITY: Osage R., presumably in Missouri. LECTOTYPE:
UMMZ 174594 (65.0), George Stolley, 1854. Gilbert (1964:129) designated lectotype. LECTO-
PARATYPES: MCZ 1914 (9), UMMZ 86915 (ex MCZ 1914) (3), USNM 120243 (ex MCZ 1914) (3),
paratopotypes.
REMARKs.-Many early literature records of this species actually refer to the closely related
N. pilsbryi, and Hubbs and Ortenburger (1929b:81-82) considered the latter to be a junior syn-
onym of N. zonatus. Hubbs and Moore (1940) later regarded the two as subspecies, and they
were so considered until I (Gilbert 1964:129-136) showed them to be specifically distinct. Bailey
et al. (1970:23) regarded Putnam as the describer of N. zonatus, but I credit this to Agassiz for
reasons discussed in my 1964 paper. Original description date 1 March 1863.
= Notropis zonatus (Agassiz)
Luxilus zonistius Jordan 1880
Jordan 1880b:239-240. TYPE LOCALITY: Suwannee (= Suwanee) Cr. (trib. to Chattahoochee
R.), ca 8 mi NW of Lawrenceville, Gwinnett County, Georgia. HOLOTYPE: USNM 23452 (68.5),
D. S. Jordan and C. H. Gilbert, summer 1876. PARATYPES: USNM 31110 (1), CAS-SU 1998 (1),
MCZ 24395 (1), BMNH 1880.1.21.55 (1), paratopotypes.
REMARKS.-Gilbert (1964:126-129) reviewed species. Jordan (1880b:240) clearly designated a
holotype in the original description, despite B6hlke's (1953:33) statement to the contrary. Origi-
nal description date 3 February 1880.
= Notropis zonistius (Jordan)
ACCOUNTS OF SPECIES PRESENTLY REFERRED TO GENUS Hybopsis, BUT WHICH MAY
ULTIMATELY BE INCLUDED IN Notropis
Rutilus amblops Rafinesque 1820
Rafinesque 1820b:51. TYPE LOCALITY: Falls of the Ohio R., at Louisville, Kentucky. TYPES:
None located.
REMARKs.-Jordan (1876a:94) initially regarded this form as of uncertain identity, but he
later (1877c:328-329) considered it to be the species with the currently-used vernacular name
"bigeye chub." The name amblops has been applied to this species ever since, despite the fact
that it was said by Rafinesque to have 10 rays (rather than 8) in both the dorsal and anal fins.
Clemmer (1970) reviewed species. Original description date (31) May 1820.
= Hybopsis amblops (Rafinesque)
Hybopsis gracilis Agassiz 1854
Agassiz 1854:358. TYPE LOCALITY: Huntsville, Alabama. SYNTYPES: None located.
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
REMARKS.-Jordan (1885b:816) was first to synonymize this form with the species amblops,
at the same time pointing out that Hyhopsis was the oldest available generic name for this and
related species. Original description date (31 December) 1854.
= Hybopsis amblops (Rafinesque)
Ceratichthys hyalinus Cope 1868
Cope, in Giinther 1868:179-180. TYPE LOCALITY: tribes. of Holston River, Virginia. SYNTYPES:
ANSP 1925-1995 (70, 37.5-65.0), USNM 118112 (formerly ANSP 1996-2000) (5, 44.5-64.0), E. D.
Cope, October 1867. Three other specimens listed in original description from BMNH.
REMARKS.-Jordan (1876a:31-32) tentatively synonymized this form with Nocomis (= Hybop-
sis) amblops, and he later (1877c:328-329) positively synonymized it with the same species. Orig-
inal description date 2 December 1868.
= Hybopsis amblops (Rafinesque)
Ceratichthys hypsinotus Cope 1870
Cope 1870:458-459. TYPE LOCALITY: Trib. to Yadkin R., Roane (= Rowan) County, North
Carolina. Type locality herein restricted by Jenkins and Lachner's designation of lectotype.
Other syntypes are from tributaries of Catawba River, McDowell County, North Carolina.
LECTOTYPE: ANSP 2005 (55.1), E. D. Cope, Autumn 1869. Lectotype herein designated by R. E.
Jenkins and E. A. Lachner. LECTOPARATYPES: ANSP 2006-2021 (16), USNM 118113 (formerly
ANSP 2024-2025) (2), paratopotypes; ANSP 2033-2035 (4 probably).
REMARKS.-One specimen each of Nocomis leptocephalus and Moxostoma papillosum re-
moved from ANSP syntypic series (originally encompassing nos. 2005-2026). One specimen of
Nocomis leptocephalus removed from USNM 118113. ANSP 2033-2035 includes one whole speci-
men, two heads without bodies, and three bodies without heads. Jenkins and Lachner (unpub.
ms.) are reviewing species. Original description date 21 November 1870.
= Hybopsis hypsinotus (Cope)
Ceratichthys labrosus Cope 1870
Cope 1870:458. TYPE LOCALITY: Upper waters of Catawba R. (Santee R. system), McDowell
and Burke counties, North Carolina. LECTOTYPE: ANSP 2045 (46.0), E. D. Cope, Autumn 1869.
Lectotype herein designated by R. E. Jenkins and E. A. Lachner. LECTOPARATYPE: ANSP 2046
(1), paratopotype. Remainder of original syntypic series (ANSP 2047-2054), comprising eight
specimens, has been reidentified by Jenkins and Lachner as Hybopsis zanema (R. E. Jenkins in
litt.). A series of four specimens (ANSP 2041-2044) probably are original syntypes, but inasmuch
as this cannot be positively established, they are not so regarded. Of these, three (nos. 2041-2043)
are the species labrosa, as here defined, and the fourth (no. 2044) has been identified as H.
zanemna.
REMARKS.-As indicated above, the syntypic series comprises two valid species. These spe-
cies, which are very similar in appearance, occur sympatrically over much of their respective
ranges, and have long been collectively called H. labrosa. One of these (the more lightly colored
of the two forms) was subsequently described (Jordan and Brayton 1878:24-25) as Ceratichthys
zanemus, which in turn has usually been regarded since as a junior synonym of H. labrosa. Cope's
description of C. labrosus could apply to either form. In the interest of nomenclatural conser-
vation, Jenkins and Lachner will call the light form H. zanema and the dark form H. labrosa.
H. labrosa and H. zanenu are most closely related to species of the subgenus Cyprinella
(genus Notropis), from which they differ in the presence of a barbel at each corer of the mouth.
Counts for lectotype: lateral-line scales 36; total vertebrae 38. Original description date 21 No-
vember 1870.
= Hybopsis labrosa (Cope)
Hyhopsis lineapunctata Clemmer and Suttkus 1971
Clemmer and Suttkus 1971:21-30, figs. 1-3. TYPE LOCALITY: Enitachope Cr., trib. to Hilla-
BULLETIN FLORIDA STATE MUSEUM
bee Cr., 2.9 mi SW of Ashland, on St. Rt. 9, Clay County, Alabama. HOLOTYPE: TU 53405 (61.0),
R. D. Suttkus and G. H. Clemmer, 13 April 1966 (field no. RDS 3877). PARATYPES: TU 40643
(46), paratopotypes; TU 12085 (8), TU 15281 (156), TU 29884 (1), TU 32729 (8), TU 40668 (12),
TU 41118 (28), TU 41132 (1), CU 51708 (6), UMMZ 168670 (3), UMMZ 168762 (5), UMMZ
168767 (1), UMMZ 175795 (2), UMMZ 177751 (19), UAIC 1036 (10), UAIC 1039 (15), UAIC
1064 (1), UAIC 1066 (26), UAIC 1067 (2), UAIC 1068 (41), UAIC 1069 (2), UAIC 1098 (13), UAIC
1247 (3), UAIC 1248 (4), UAIC 1249 (2), UAIC 1250 (?), UAIC 1251 (2), UAIC 1282 (20), UAIC
1284 (5), UAIC 1308 (1), UAIC 1309 (1), UAIC 1310 (2), UAIC 1316 (3), UAIC 1317 (6), UAIC
1318 (6), UAIC 1319 (3), UAIC 1320 (3), UAIC 1362 (4), UAIC 1375 (?), UAIC 1376 (?), UAIC
1377 (6), UAIC 1378 (25), UAIC 1379 (3), UAIC 1380 (1), UAIC 1381 (5), UAIC 1382 (7), UAIC
1383 (2), UAIC 1487 (2), UAIC 1497 (17), UAIC 1504 (3), UAIC 1505 (2), UAIC 1509 (2), UAIC
1517 (9), UAIC 1519 (14), UAIC 1522 (92), UAIC 1523 (1), UAIC 1529 (5), UAIC 1911 (2).
REMARKS.-Species closely related to H. amblops. Original description date 30 September
1971.
= Hybopsis lineapunctata Clemmer and Suttkus 1971
Nocomis ruhrifrons Jordan 1877
Jordan 1877c:330. TYPE LOCALITY: South Fork of Ocmulgee R., at Flat Rock, DeKalb County,
Georgia. SYNTYPES: USNM 17863 (7, 45.3-55.4), USNM 20146 (1, 59.3), MCZ 24381 (1, 57.0),
D. S. Jordan and C. H. Gilbert, summer 1876.
REMARKs.-Species closely related to H. amblops. Inasmuch as the name rubrifrons is preoccu-
pied in Notropis by Alburnus rubrifrons Cope 1865, a new name will be required for this species
if and when it is eventually transferred to Notropis. Clemmer (1970) reviewed species. Original
description date (30) June 1877.
= Hybopsis rubrifrons (Jordan)
Hybopsis winchelli Girard 1857
Girard 1857:211. TYPE LOCALITY: Black Warrior R., at Selma, Alabama, SYNTYPE: USNM 2
(one pair of pharyngeal teeth in osteological collection), Prof. A. Winchell.
REMARKS.-Jordan (1885a:127) regarded this species as a synonym of H. amblops. It has either
been so considered since (Jordan and Evermann 1896:321; Bailey, Winn and Smith 1954:124) or
treated as a subspecies of H. amblops (Hubbs 1951a:5). Recent studies by Clemmer (1970) indi-
cate full species status. Counts: pharyngeal teeth 1,4-4,1. Original description date 25 April 1857.
= Hybopsis winchelli Girard
Ceratichthys zanemus Jordan and Brayton 1878
Jordan and Brayton 1878:24-25. TYPE LOCALITY: Saluda R., at Farr's Mills, W of Greenville,
Greenville County, South Carolina. LECTOTYPE: USNM 31126 (34.0), D. S. Jordan, A. W. Bray-
ton, C. H. Gilbert et al., summer 1877. Lectotype herein designated by R. E. Jenkins and E. A.
Lachner.
REMARKS.-Type in very poor condition, with fins mostly broken, body almost completely de-
scaled, and head almost separated from body. Counts: pharyngeal teeth 4,1, with major teeth
well hooked; total vertebrae 39. Although only one type apparently exists, it is not regarded as
the holotype because Jordan and Brayton, in the original description, mentioned having col-
lected a number of specimens. For further remarks, see account of Ceratichthys labrosus. Origi-
nal description date (31 December) 1878.
= Hybopsis zanema (Jordan and Brayton)
Vol. 23, No. 1
GILBERT: NOTROPIS TYPE CATALOGUE
LITERATURE CITED
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Vol. 23, No. 1
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Vol. 23, No. 1
1978
GILBERT: NOTROPIS TYPE CATALOGUE
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